Family DORADIDAE Bleeker 1858 (Thorny Catfishes)

Updated 12 Aug. 2024
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Subfamily ACANTHODORADINAE Sabaj & Mariangeles 2021

Acanthodoras Bleeker 1862 acanthus (L.), from ákantha (Gr. ἄκανθα), thorn, probably referring to upper and hind parts of body of A. cataphractus covered by bony thorn-bearing; Doras, type genus of family

Acanthodoras cataphractus (Linnaeus 1758) katáphraktos (Gr. κατάφρακτος), clad in armor, referring to any or all of the following: heavily ossified nuchal shield; bony spine-like posterior cleithral (humeral) process; sturdy and well-serrated pectoral-fin spines; deep, bony thorn-bearing plates along sides of body from tympanal region to caudal-fin base

Acanthodoras depressus (Steindachner 1881) Latin for pressed down, referring to strongly depressed head, “almost completely flat across the top” (translation)

Acanthodoras spinosissimus (Eigenmann & Eigenmann 1888) Latin for very spiny or thorny, presumably referring to “short sharp” spines on humeral process, “a series of which near the lower margin, is enlarged”


Spotted Thorny Cats
Subfamily ASTRODORADINAE Higuchi, Birindelli, Sousa & Britski 2007

Amblydoras Bleeker 1862 amblýs (Gr. ἀμβλύς), blunt, allusion not explained, perhaps referring to smooth (non-serrated) dorsal-fin spine of A. affinis; Doras, type genus of family

Amblydoras affinis (Kner 1855) Latin for related, referring to close similarity to Doras dentatus (=Platydoras costatus), its presumed congener at the time

Amblydoras gonzalezi (Fernández-Yépez 1968) in honor of Venezuelan civil engineer Marcelo González Molina (1923–2000), who provided access to the type locality

Amblydoras monitor (Cope 1872) allusion not explained; in presenting his paper to the Academy of Natural Sciences of Philadelphia in 1871, Cope reportedly described the swim bladder of A. monitor as having a “gun-boat style of shape,” perhaps a reference to USS Monitor, an ironclad warship commissioned in 1862 during the U.S. Civil War that received much attention at the time (name could also be comparing the fish’s bony shields to the Monitor’s ironclad hull; note that in the same paper Cope compared Physopyxis lyra to a “miniature iron-clad with mast and outriggers”)

Amblydoras nauticus (Cope 1874) icus (L.), belonging to: Nauta, Peru, type locality

Amblydoras nheco (Higuchi, Birindelli, Sousa & Britski 2007) named for town of Nhecolândia, Rio Paraguay basin, Brazil, type locality; nheco apparently does not have any particular significance other than having probably been a nickname for the town founder

Anadoras Eigenmann 1925 án (Gr. privative, ἄν), i.e., without, referring to absence of plates between dorsal and adipose fins; Doras, type genus of family

Anadoras grypus (Cope 1872) from grypós (Gr. γρυπός), curved or hook-nosed, referring to its short, smoothly curved snout (Mark H. Sabaj Pérez, pers. comm.)

Anadoras insculptus (Miranda Ribeiro 1912) Latin for engraved with markings, i.e., sculpted, referring to sculptured nature of exposed bones of cranium and pectoral girdle (Mark H. Sabaj Pérez, pers. comm.)

Anadoras regani (Steindachner 1908) patronym not identified but almost certainly in honor of English ichthyologist Charles Tate Regan (1878–1943), Natural History Museum (London)

Anadoras weddellii (Castelnau 1855) in honor of British physician-botanist Hugh Algernon Weddell (1819–1877), who presented holotype (a dried skin) and a drawing of it

Astrodoras Bleeker 1862 astro, from astḗr (Gr. ἀστήρ), star, from the specific epithet asterifrons, presumably referring to its elevated (i.e., stargazing) eyes; Doras, type genus of family

Astrodoras asterifrons (Kner 1853) asterías (Gr. ἀστερίας), a starred or starry fish or bird; frons (L.), face or brow, allusion not explained, presumably referring to raised superciliary margin, creating elevated (e.g., stargazing) eyes

Hypodoras Eigenmann 1925 hypó (Gr. ὑπό), under or beneath, allusion not explained, perhaps referring to anterior two thirds of adipose fin under a rhomboidal plate; Doras, type genus of family

Hypodoras forficulatus Eigenmann 1925 Latin for forked, referring to posterior swim bladder, described as “forked (divided into two horns)”

Physopyxis Cope 1871 physo-, bladder; pyxis, box, referring to “osseus box” that encloses and protects large, greatly developed swim bladder of P. lyra

Physopyxis ananas Sousa & Rapp Py-Daniel 2005 Ananas, pineapple genus (Bromeliaceae), referring to stout body and spiny (i.e., pineapple-like) appearance due to additional row(s) of spines on lateral plates

Physopyxis cristata Sousa & Rapp Py-Daniel 2005 Latin for crested, referring to externally visible series of emergent neural spines

Physopyxis lyra Cope 1872 Latin for lyre, referring to the postcoracoid processes, “curved, and the extremities dilated outwards, so as to present with the thoracic portion exactly the form of the Grecian lyre”

Scorpiodoras Eigenmann 1925 scorpio (L.), from skorpiós (Gr. σκορπιός), scorpion, referring to “banjo- or scorpion-shaped” posterior swim bladder of S. heckelii; Doras, type genus of family

Scorpiodoras bolivarensis (Fernández-Yépez 1968) ensis, Latin suffix denoting place: Estado Bolívar, Venezuela, type locality

Scorpiodoras heckelii (Kner 1855) in honor of Austrian ichthyologist Johann Jakob Heckel (1790–1857), who provided manuscript names and descriptions for many of the doradids that Kner ultimately described

Scorpiodoras liophysus Sousa & Birindelli 2011 lio, from leí̄os (Gr. λεῖος), smooth or bald; physus, from physa (Gr. φύσα), bladder, referring to absence of a secondary swim bladder

Eastern Brazil Thorny Cats
Subfamily WERTHEIMERINAE Birindelli 2014

Franciscodoras Eigenmann 1925 Francisco, named for rio São Francisco basin, Brazil, where F. marmoratus is endemic; Doras, type genus of family

Franciscodoras marmoratus (Lütken 1874) Latin for marbled, referring to its coloration

Kalyptodoras Higuchi, Britski & Garavello 1990 kalýpto (Gr. καλύπτω), cover, referring to embedded condition of most lateral scutes; Doras, type genus of family

Kalyptodoras bahiensis Higuchi, Britski & Garavello 1990 ensis, Latin suffix denoting place: Bahia State, Brazil, where type locality (rio Paraguaçu) is situated

Wertheimeria Steindachner 1877ia (L. suffix), belonging to: Mr. Wertheimer, who collected holotype, either Louis Wertheimer of the Thayer Expedition to Brazil (1865–1866), or Achilles Wertheimer, who died on the expedition due to a snakebite (see Pogonopoma wertheimeri, Loricariidae: Rhinelepinae)

Wertheimeria maculata Steindachner 1877 Latin for spotted, referring to large, oval, blue-gray spots on upper body, top of head, and pectoral and dorsal fins


Subfamily AGAMYXINAE Sabaj & Mariangeles 2021

Agamyxis Cope 1878 aga-, from ágan (Gr. ἄγαν), much or very; myxis, from mýxa (Gr. μύξα), slime, allusion not explained; per Mark H. Sabaj Pérez (pers. comm.), Agamyxis closely resembles and sometimes occurs with Acanthodoras, which, when disturbed, exude a white milky mucous from the axillary region of their pectoral spines; perhaps John Hauxwell, who collected holotype of A. pectinifrons (type species of Agamyxis) shared such an observation with Cope

Agamyxis albomaculatus (Peters 1877) albus (L.), white; maculatus (L.), spotted, referring to white spots on body, abdomen and caudal fin

Agamyxis pectinifrons (Cope 1870) pectinatus (L.), raked or combed; frons (L.), brow or forehead, referring to strongly toothed preorbital bone (or “comb”), the first bone in the infraorbital series (also known as the lacrimal), forming a serrated crest anterior to the orbit (Mark H. Sabaj Pérez, pers. comm.)


Subfamily RHINODORADINAE Sabaj & Mariangeles 2021

Rhinodoras Bleeker 1862 rhinos (Gr. ῥινός), genitive of rhís (ῥίς), snout, referring to pointed snout of R. dorbignyi; Doras, type genus of family

Rhinodoras armbrusteri Sabaj Pérez 2008 in honor of Jonathan W. Armbruster (b. 1969), curator of Fishes, Auburn University Museum (Alabama, USA), for “sterling” contributions to the collection and study of Neotropical fishes, and for “deftly” leading the expedition to Guyana that led to the discovery of this species

Rhinodoras boehlkei Glodek, Whitmire & Orcés V. 1976 in honor of American ichthyologist James E. Böhlke (1930–1982), Academy of Natural Sciences of Philadelphia, for his work on South American freshwater fishes

Rhinodoras dorbignyi (Kner 1855) in honor of Alcide d’Orbigny (1802–1857), French explorer and naturalist, who collected in South America for the Muséum national d’Histoire naturelle (Paris) from 1827–1833

Rhinodoras gallagheri Sabaj Pérez, Taphorn & Castillo G. 2008 in honor of Francis Richard Gallagher (b. 1935), mailroom supervisor, Academy of Natural Sciences of Philadelphia (1967–2003), for dedicated service to the global community of taxonomists and systematists via the shipping and receiving of countless loans of biological specimens

Rhinodoras thomersoni Taphorn & Lilyestrom 1984 in honor of American ichthyologist Jamie E. Thomerson (1935–2015), Southern Illinois University, who introduced the senior author to the study of fishes and led his first trip to South America (he also led expedition during which holotype was collected)

Rhynchodoras Klausewitz & Rössel 1961 rhýnchos (Gr. ῥύγχος), snout, referring to its turned-down, proboscis-like snout; Doras, type genus of family

Rhynchodoras castilloi Birindelli, Sabaj Pérez & Taphorn 2007 in honor of Venezuelan biologist Otto E. Castillo G., who collected much of the type material, for his lifelong dedication to the study and stewardship of his country’s rich diversity of freshwater fishes

Rhynchodoras woodsi Glodek 1976 in honor of Loren P. Woods (1914–1979), Curator of Fishes, Field Museum of Natural History (Chicago), for numerous contributions to ichthyology

Rhynchodoras xingui Klausewitz & Rössel 1961 of the rio Xingu, Brazil, type locality

Fimbriate Barbel Thorny Cats
Subfamily DORADINAE Bleeker 1858

Anduzedoras Fernández-Yépez 1968 in honor of Pablo J. Anduze (1902–1989), Venezuelan explorer, ethnologist and medical entomologist, and later governor of Amazonas State, Venezuela, whom Fernández-Yépez said dedicated many years to practicing medicine in the jungle, and always found time to collect zoological specimens; Doras, type genus of family

Anduzedoras oxyrhynchus (Valenciennes 1821) sharp-snouted, from oxýs (Gr. ὀξύς), sharp, and rhýnchos (Gr. ῥύγχος), snout, referring to compressed head with relatively pointed snout

Centrochir Agassiz 1829 centro-, from kéntron (Gr. κέντρον), any sharp point; chir, from cheír (Gr. χείρ), hand (homologous to the pectoral fin), referring to its unrayed pectoral-fin spines

Centrochir crocodili (Humboldt 1821) of a crocodile (L. crocodilus), referring to its local name in Colombia, pexe que mata el Cayman (fish that kills caiman), referring to its sharp pectoral fins, known to kill caiman (or crocodiles) that attempt to eat it

Centrodoras Eigenmann 1925 centro-, from kéntron (Gr. κέντρον), any sharp point, allusion not explained, perhaps referring to dorsal spine of C. brachiatus, “with antrorse hooks in front, with straight to retrorse hooks behind”; Doras, type genus of family

Centrodoras brachiatus (Cope 1872) Latin for having branches like arms, presumably referring its to “very large” pectoral spines

Centrodoras hasemani (Steindachner 1915) in honor of American zoologist John D. Haseman (1882–1969), field collector for the Carnegie Museum of Natural History, who collected holotype

Doraops Schultz 1944 ṓps (Gr. ὦψ), eye or face (i.e., appearance), similar to “other members of the family Doradidae”

Doraops zuloagai Schultz 1944 in honor of Venezuelan geologist Guillermo Zuloaga (1904–1984), assistant chief of exploration, Standard Oil Co. of Venezuela, who was largely responsible for inviting Schultz to study the fishes of the Maracaibo Basin

Doras Lacepède 1803 dorás (Gr. δοράς), animal hide or skin, but said by Lacepède to mean cuirass (or breastplate), which were originally made of leather, probably referring to hard bony plates and/or thick, leathery skin (name does not mean “spear” as reported by others; that would be dóry, Gr. δόρυ)

Doras carinatus (Linnaeus 1766) Latin for keeled, referring to spines on midlateral scutes (which reminded Linnaeus of the longitudinal keels in “Scombris,” presumably Atlantic Mackerel Scomber scombrus)

Doras higuchii Sabaj Pérez & Birindelli 2008 in honor of Horácio Higuchi, Universidade de São Paulo, for his “groundbreaking contributions” to the systematics of doradid catfishes

Doras micropoeus (Eigenmann 1912) micro-, from mikrós (Gr. μικρός), small; poeus, from poiéō (Gr. ποιέω), to make or produce, referring to reduced or “rudimentary” anterior midlateral scutes

Doras phlyzakion Sabaj Pérez & Birindelli 2008 phlyzákion (Gr. φλυζάκιον), diminutive of phlýktaina (φλύκταινα), blister, i.e., a small blister, referring to abundant pores on ventral surface

Doras zuanoni Sabaj Pérez & Birindelli 2008 in honor of Brazilian ichthyologist Jansen Alfredo Sampaio Zuanon, for his “extensive and valuable” contributions to the collection, taxonomy and natural history of Neotropical fishes, including discovery of this species

Hassar Eigenmann & Eigenmann 1888 Arawak name for species of Doras in Guyana

Hassar affinis (Steindachner 1881) Latin for related, described as a variation of the closely related H. orestis

Hassar gabiru Birindelli, Fayal & Wosiacki 2011 in honor of Leandro Melo de Sousa, known to his friends as “Gabiru,” for his many contributions to the understanding of the Doradidae, including his M.Sc. and Ph.D. dissertations; he also helped collect part of the type series [a noun in apposition, without the patronymic “i”]

Hassar orestis (Steindachner 1875)is, Latin genitive singular of: American geologist Orestes Saint John (1841–1921), who collected holotype during Thayer Expedition to Brazil (1865–1866 (name was recommended by Louis Agassiz, leader of the Expedition)

Hassar shewellkeimi Sabaj Pérez & Birindelli 2013 in honor of Shewell “Bud” DeBenneville Keim (1918–2014), electrical engineer, World War II veteran, and nephew of Henry Weed Fowler (1878–1965), the first full-time curator of fishes at the Academy of Natural Sciences of Philadelphia, for his generous support of Academy ichthyology, including the preservation of Fowler’s legacy and the stewardship of his uncle’s fishes

Hassar wilderi Kindle 1895 in honor of American zoologist Burt Green Wilder (1841–1925), Cornell University, who sent the collection made by C. F. Hartt in Brazil to Carl H. Eigenmann for identification (and then transmitted to Kindle for final determination)

Hemidoras Bleeker 1858 hemi-, from hḗmisys (Gr. ἥμισυς), half, referring to species related to (and previously placed in) Doras but which lack teeth in the jaws and palate

Hemidoras boulengeri Steindachner 1915 in honor of Belgian-born British ichthyologist-herpetologist Georges A. Boulenger (1858–1937), British Museum (Natural History)

Hemidoras morei (Steindachner 1881) patronym not identified nor can identity be inferred based on available information; could this be named after the same person (also unknown) Steindachner honored with the bryconid name Brycon moorei (note spelling) in 1878?

Hemidoras morrisi Eigenmann 1925 in honor of Percival Morris, who collected holotype and served as Eigenmann’s assistant, majordomo and interpreter in the Iquitos region of Peru

Hemidoras stenopeltis (Kner 1855) sténos (Gr. στένος), narrow; peltis, from péltē (Gr. πέλτη), small shield, probably referring to long and narrow humeral process

Hemidoras stuebelii (Steindachner 1882) in honor of German geologist-vulcanologist Alphons Stübel (1835–1904), who collected holotype

Leptodoras Boulenger 1898 leptós (Gr. λεπτός), thin or slender, referring to longer bodies (with longer anal fins) of L. acipenserinus and L. juruensis compared with Oxydoras; Doras, type genus of family

Leptodoras acipenserinus (Günther 1868)inus (L.), adjectival suffix: Acipenser, sturgeon genus, i.e., sturgeon-like, presumably referring to “peculiar” shape of head and snout, “elongate triangular, pointed, and much depressed in its anterior portion”

Leptodoras cataniai Sabaj Pérez 2005 in honor of David Catania, for his dedicated service to the ichthyological community since 1985 as Collection Manager of Fishes, California Academy of Sciences

Leptodoras copei (Fernández-Yépez 1968) in honor of American zoologist-paleontologist Edward Drinker Cope (1840–1897), for many “excellent” contributions to the study of Neotropical fishes, including doradid catfishes

Leptodoras hasemani (Steindachner 1915) in honor of American zoologist John D. Haseman (1882–1969), Carl Eigenmann’s student and field collector for the Carnegie Museum of Natural History, who collected holotype

Leptodoras juruensis Boulenger 1898 ensis, Latin suffix denoting place: río Jurua, Brazil, type locality

Leptodoras linnelli Eigenmann 1912 in honor of George Linnell, Essequibo Exploring Company, who provided Eigenmann with boats and crewmen for his expedition to British Guiana

Leptodoras marki Birindelli & Sousa 2010 in honor of Mark Sabaj Pérez (b. 1969), Collection Manager of Fishes at the Illinois Natural History Survey (1995–2000) and the Academy of Natural Sciences of Philadelphia (2000–present), for dedicated service to the ichthyological community, and for encouraging the study of doradid catfishes as a co-principal investigator of the All Catfish Species Inventory

Leptodoras myersi Böhlke 1970 in honor of Böhlke’s professor and good friend George S. Myers (1905–1985), ichthyologist, Stanford University

Leptodoras nelsoni Sabaj Pérez 2005 in honor of Douglas W. Nelson, for “dedicated” service to the ichthyological community since 1993 as Collection Manager of Fishes, University of Michigan Museum of Zoology

Leptodoras oyakawai Birindelli, Sousa & Sabaj Pérez 2008 in honor of Osvaldo Takeshi Oyakawa, for dedicated service to the ichthyological community since 1989 as Collection Manager of the Fish Collection at the Museu de Zoologia da Universidade de São Paulo

Leptodoras praelongus (Myers & Weitzman 1956) prae– (L.), in front of; longus (L.), long, presumably referring to longer snout compared with Hassar lipophthalmus (=Anduzedoras oxyrhynchus)

Leptodoras rogersae Sabaj Pérez 2005 in honor of Mary Ann Rogers, for her dedicated service to the ichthyological community since 1988 as Collection Manager of Fishes, Field Museum of Natural History (Chicago)

Lithodoras Bleeker 1862 líthos (Gr. λῐ́θος), stone, presumably referring to heavily armored body, including belly; Doras, type genus of family

Lithodoras dorsalis (Valenciennes 1840) Latin for of the back, presumably referring to 5–6 oblong plates between dorsal and adipose fins

Megalodoras Eigenmann 1925 megalo-, from mégas (Gr. μέγας), big, presumably referring to large size of M. uranoscopus (up to 60 cm SL and 4.8 kg); Doras, type genus of family

Megalodoras guayoensis (Fernández-Yépez 1968) ensis, Latin suffix denoting place: Caño de Guayo, Delta Amacuro, Estado Delta, Venezuela, type locality

Megalodoras uranoscopus (Eigenmann & Eigenmann 1888) ouranós (Gr. οὐρανός), heaven or sky; skopós (Gr. σκοπός), looker, contemplator or viewer, “eye more superior [on top of head] than lateral [on sides]”

Nemadoras Eigenmann 1925 nḗma (Gr. νῆμα), thread, referring to “simple” (vs. fringed) maxillary barbels; Doras, type genus of family

Nemadoras elongatus (Boulenger 1898) Latin for prolonged, referring to elongate body (depth 5 times in TL) compared with presumed congeners at the time

Nemadoras hemipeltis (Eigenmann 1925) hemi-, from hḗmisys (Gr. ἥμισυς), half; peltis, from péltē (Gr. πέλτη), small shield, allusion not explained, perhaps referring to fontanel “not continued as a groove to the dorsal plate”

Nemadoras humeralis (Kner 1855) Latin for humeral, presumably referring to rough and very wide humeral process, its greatest width equal to the eye

Orinocodoras Myers 1927 Orinoco, referring to Orinoco River basin, Venezuela, type locality of O. eigenmanni; Doras, type genus of family

Orinocodoras eigenmanni Myers 1927 in honor of German-born American ichthyologist Carl H. Eigenmann (1863–1927), “who recently [1925] placed the classification of the Doradidae on a firm foundation”

Ossancora Sabaj Pérez & Birindelli 2011 oss (L.), bone; ancora (L.), anchor, referring to shape and articulation of pectoral spine and posterior cleithral and coracoid processes, which resemble shank and flukes, respectively, of a Danforth anchor

Ossancora asterophysa Birindelli & Sabaj Pérez 2011 aster, star; physa, bladder, referring to proliferation of diverticula along periphery of swim bladder

Ossancora eigenmanni (Boulenger 1895) in honor of German-born American ichthyologist Carl H. Eigenmann (1863–1927), author of an “excellent” synopsis (1890) of South American catfishes

Ossancora fimbriata (Kner 1855) Latin for fringed, probably referring to fimbriae present on maxillary and mental barbels (may also refer to many diverticula on swim bladder)

Ossancora punctata (Kner 1855) Latin for spotted, presumably referring to dark dots conspicuously evident on head and body of syntypes

Oxydoras Kner 1855 oxýs (Gr. ὀξύς), sharp or pointed, allusion not explained, perhaps referring to “conically elongated” (translation) snouts of O. kneri (reported as O. niger) and O. (=Hemidoras) stenopeltis; Doras, type genus of family

Oxydoras kneri Bleeker 1862 in honor of Austrian ichthyologist Rudolph Kner (1810–1869), who described this catfish under the name Doras (Oxydoras) niger in 1855

Oxydoras niger (Valenciennes 1821) Latin for dark or black, presumably referring to its dark-brown or black coloration

Oxydoras sifontesi Fernández-Yépez 1968 in memory of Venezuelan meteorologist-hydrologist Ernesto Sifontes (1881–1959), who devoted much of his life to studying the Río Orinoco, where this catfish occurs

Platydoras Bleeker 1862 platýs (Gr. πλατύς), flat or broad, presumably referring to depressed head; Doras, type genus of family

Platydoras armatulus (Valenciennes 1840) diminutive of armatus (L.), armed with a weapon, from armadillo, Spanish for “little armored one,” apparently its local name in Brazil, probably referring to bony shields along middle of body, covered with backwards-pointing spines

Platydoras birindellii Sousa, Chaves, Akama, Zuanon & Sabaj 2018 in honor of colleague and friend José Luís O. Birindelli (b. 1979), Universidade de São Paulo, “for enriching our knowledge of doradid catfishes and inspiring the next generation of Neotropical ichthyologists with his enthusiasm and integrity”

Platydoras brachylecis Piorski, Garavello, Arce H. & Sabaj Pérez 2008 brachýs (Gr. βραχύς), short; lekís (Gr. λεκίς) or lékos (λέκος) dish, pot or pan (authors say plate or dish), referring to its relatively shallow midlateral scutes

Platydoras costatus (Linnaeus 1758) Latin for ribbed, possibly referring to rib-like appearance of longitudinal bony plates

Platydoras hancockii (Valenciennes 1840) in honor of British naturalist (and father of modern taxidermy) John Hancock (1808–1890); in 1829 he reported how this catfish, under the name Doras (now Platydoras) costatus, can crawl on land during droughts in search of water

Pterodoras Bleeker 1862 pterus, from pterón (Gr. πτερόν) or ptéryx (πτέρυξ), fin, referring to forked caudal fin; Doras, type genus of family

Pterodoras granulosus (Valenciennes 1821) granum (L.), seed or grain; –osus (L.), suffix connoting fullness, presumably referring to 23–28 shallow lateral plates (or scutes) along length of body, which may give the scaleless skin a granulated appearance

Pterodoras rivasi (Fernández-Yépez 1950) in honor of Luis A. Rivas L. (no other information available), who collected fishes with Fernández-Yépez and assisted with the description of this species

Tenellus Birindelli 2014 Latin for somewhat tender or delicate, referring to delicate appearance of all included species

Tenellus cristinae (Sabaj Pérez, Arce H., Sousa & Birindelli 2014) in honor of Maria Cristina Sabaj Pérez, teacher at Friends’ Central School (Wynnewood, Pennsylvania, USA), for her contributions to the collection of the type series and to the well-being of the senior author (her husband)

Tenellus leporhinus (Eigenmann 1912) lepo-, from lepis (L.), hare or rabbit; rhinós (Gr. ῥινός), genitive of rhís (ῥίς), nose, referring to its “peculiar leporine snout”

Tenellus ternetzi (Eigenmann 1925) in honor of Swiss-born ichthyologist and naturalist Carl Ternetz (1870–1928), who collected holotype

Tenellus trimaculatus (Boulenger 1898) tri– (L.), three; maculatus (L.), spotted, referring to black spot on dorsal fin and at base of each caudal-fin lobe

Trachydoras Eigenmann 1925 trachýs (Gr. τραχύς), jagged or rough, referring to granular opercle, preopercle and coracoid-process; Doras, type genus of family

Trachydoras brevis (Kner 1853) Latin for short, referring to its short (length) but high (height) body

Trachydoras gepharti Sabaj & Arce H. 2017 in honor of George W. Gephart, Jr., President & CEO of the Academy of Natural Sciences of Drexel University from 2010 to 2017, for his “bold, deft and heartfelt leadership of a Glorious Enterprise into its third century” [“Glorious Enterprise” alludes to the title of a 2012 book about the Academy, which is America’s oldest natural history museum]

Trachydoras microstomus (Eigenmann 1912) small-mouthed, from mikrós (Gr. μικρός), small, and stóma (Gr. στόμα), mouth, its width “equal to half the distance between gill-openings”

Trachydoras nattereri (Steindachner 1881) patronym not identified but probably in honor of Austrian naturalist Johann Natterer (1787–1843), who explored South America and collected specimens for 18 years

Trachydoras paraguayensis (Eigenmann & Ward 1907)ensis, Latin suffix denoting place: Corumba, Paraguay, type locality

Trachydoras steindachneri (Perugia 1897) patronym not identified but clearly in honor of Austrian ichthyologist Franz Steindachner (1834–1919), who described several doradid taxa