Order ACANTHURIFORMES (part 1): Families LOBOTIDAE, POMACANTHIDAE, DREPANEIDAE and CHAETODONTIDAE

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v. 4.0 – 10 Feb. 2023  view/download PDF

Family LOBOTIDAE Tripletails and Tigerperches
3 genera · 15 species · Taxonomic note: contains taxa placed in the families Datnioididae and Hapalogenyidae by some workers.

Datnioides Bleeker 1853    oides, having the form of: Datnia (=Mesopristes, Centrarchiformes: Terapontidae), then classified together in the catch-all family Percidae (perches) and presumed to be related because of their lack of palatine teeth

Datnioides campbelli Whitley 1939    in honor of Flight-Lieutenant Stuart Campbell (1903-1988), who collected type

Datnioides microlepis Bleeker 1854    micro-, small; lepis, scale, referring to small scales compared to D. polota

Datnioides polota (Hamilton 1822)    presumably local Gangetic name for this species in India

Datnioides pulcher (Kottelat 1998)    beautiful, allusion not explained, perhaps referring to barred color pattern

Datnioides undecimradiatus (Roberts & Kottelat 1994)    undecim, eleven; radiatus, rayed, referring to “most frequent count” of soft anal-fin rays

Hapalogenys Richardson 1844    hapalos, soft; genys, chin, referring to “velvety softness of the chin and lower lip, which is made more conspicuous by contrast with the rigidly rough scales that cover the rest of the head”

Hapalogenys analis Richardson 1845    anal, referring to “very strong, deeply striated second anal spine taller than the soft part of the fin,” presumably stouter and longer than anal-fin spine of H. nitens (=nigripinnis)

Hapalogenys bengalensis Mohapatra, Ray & Kumar 2013    ensis, suffix denoting place: northern Bay of Bengal, West Bengal coast of India, type locality                     

Hapalogenys dampieriensis Iwatsuki & Russell 2006    ensis, suffix denoting place: Dampierian Province, a biogeographic region extending from approximately Geraldton in Western Australia across northern Australia to Cape York, approximating the distribution of this species

Hapalogenys filamentosus Iwatsuki & Russell 2006    filamentous, referring to filamentous first ray of pelvic fin

Hapalogenys kishinouyei Smith & Pope 1906    in honor of Kamakichi Kishinouye (1867-1929), head of the Imperial Fisheries Bureau of Japan

Hapalogenys merguiensis Iwatsuki, Satapoomin & Amaoka 2000    ensis, suffix denoting place: Mergui Basin, southern Myanmar Sea, type locality                       

Hapalogenys nigripinnis (Temminck & Schlegel 1843)    nigri-, black; pinnis, fin, referring to blackish fins

Hapalogenys sennin Iwatsuki & Nakabo 2005    Japanese term for a djinn, sage or hermit, often depicted in Japanese art with a beard, referring to this fish’s barbels on lower lip and solitary habitat, commonly observed in small caves, holes, and under over-hanging rocks

Lobotes Cuvier 1829    lobed, referring to how soft parts of dorsal, anal and caudal fins appear to resemble one three-lobed fin (hence the name tripletail)

Lobotes pacifica Gilbert 1898    icus, belonging to: Pacific Ocean, referring to distribution in eastern Pacific from southern California (USA) to Peru, including Gulf of California

Lobotes surinamensis (Bloch 1790)    ensis, suffix denoting place: Caribbean Sea off Suriname, type locality (but circumglobal in tropical and warm temperate seas except for eastern Pacific)


Family POMACANTHIDAE Angelfishes
13 genera/subgenera · 90 species

Apolemichthys Burton 1934    etymology not explained, perhaps a-, without and polemos, war, i.e., unfit for battle, proposed as a subgenus of Holacanthus without “powerful” preorbital spines; ichthys, fish (name proposed by Fraser-Brunner in 1933 but unavailable since he included multiple species without designating which one was the type; authorship is credited to Maurice Burton, a compiler for the Zoological Record, who designated type) [nests within Centropyge according to some workers]

Apolemichthys arcuatus (Gray 1831)    bowed or arched, referring to a “broad black arched band from the eye to the caudal end of the dorsal fin”                        

Apolemichthys griffisi (Carlson & Taylor 1981)    in honor of Nixon Griffis (1917-1993), Trustee of New York Zoological Society and Patron of the New York Aquarium, whose “interest in marine life and who generosity resulted in [the authors’] collecting trip to Canton Island [northern Phoenix Islands, South Pacific] and the discovery of this new species”

Apolemichthys guezei (Randall & Maugé 1978)    in honor of marine biologist Paul Guézé, who collected type specimens and made them available for study                         

Apolemichthys kingi Heemstra 1984    in honor of Dennis King (Durban, South Africa), structural engineer and underwater photographer, who discovered this angelfish and collected type

Apolemichthys trimaculatus (Cuvier 1831)    tri-, three; maculatus, spotted, referring to black spot on forehead and one indistinct spot on each side of head behind gill cover of adults

Apolemichthys xanthopunctatus Burgess 1973    xanthos, yellow; punctatus, spotted, referring to golden scale centers, giving it a spotted appearance                                   

Apolemichthys xanthotis (Fraser-Brunner 1950)    xanthos, yellow; otis, ear, referring to “brilliant lemon yellow spot (white in spirits) on external face of post-temporal, above gill-cover”

Apolemichthys xanthurus (Bennett 1833)    xanthus, yellow; oura, tail, referring to yellow caudal fin                   

Centropyge Kaup 1860    kentron, thorn or spine; pyge, rump or rear, referring to four (actually three) anal-fin spines of C. tibicen

Subgenus Centropyge

Centropyge aurantia Randall & Wass 1974    orange, referring to orange (or reddish-orange) ground color

Centropyge bicolor (Bloch 1787)    bi-, two, referring to two colors, yellow in the front (with blue around eyes) and blue in the back

Centropyge bispinosa (Günther 1860)    bi-, two; spinosa, spined, referring to two spines at angle of preoperculum

Centropyge cocosensis Shen, Chang, Delrieu-Trottin & Borsa 2016    –ensis, suffix denoting place: Cocos (Keeling) Islands, east Indian Ocean, type locality (also occurs at Christmas Island) [possibly intermediate between sympatric color morphs corresponding to C. eibli and C. vrolikii]

Centropyge colini Smith-Vaniz & Randall 1974    in honor of Patrick L. Colin (b. 1946), then at the University of Puerto Rico (now at the Coral Reef Research Foundation, Koror, Palau), for his contributions to the success of the expedition during which types of this species and C. joculator were collected (Colin helped collect them both)

Centropyge debelius Pyle 1990    in honor of underwater photographer and author Helmut Debelius (b. 1947), who presented an underwater photograph of this angelfish to Pyle and was the first to sight the holotype [presumably a noun in apposition, without the patronymic “i”]

Centropyge eibli Klausewitz 1963    in honor of ethologist and part-time ichthyologist Irenäus Eibl-Eibesfeldt (1928-2018); he collected three of the type specimens as part of the Xarifa Expedition to the Indian Ocean

Centropyge ferrugata Randall & Burgess 1972    iron-red, i.e., rusty, referring to body color (which varies from pale cream to bright red)

Centropyge flavipectoralis Randall & Klausewitz 1977    flavus, yellow; pectoralis, pectoral, referring to yellow pectoral fins, which starkly contrast against the fish’s dark body

Centropyge flavissima (Cuvier 1831)    very yellow, referring to color of body and fins

Centropyge heraldi Woods & Schultz 1953    in honor of Earl S. Herald (1914-1973), ichthyologist and Director of the Steinhart Aquarium in San Francisco (1952-1966), who helped collect type

Centropyge hotumatua Randall & Caldwell 1973    named for Hotumatua, legendary Polynesian chieftain who first colonized Easter Island (type locality) [presumably a noun in apposition, without the patronymic “i”]

Centropyge interrupta (Tanaka 1918)    interrupted, allusion not explained, presumably referring to blackish brown bars on head, “which are interrupted here and there, forming elongate spots” (translation)

Centropyge joculator Smith-Vaniz & Randall 1974    jester or clown, referring to bold and gaudy coloration in life

Centropyge loriculus (Günther 1874)    etymology not explained, perhaps diminutive of lorius, parrot, source of the hanging parrot genus Loriculus (if so, allusion not evident), or perhaps (per Mazza 2013, Monaco Nature Encyclopedia) diminutive of lorica, breastplate or corset, referring to black vertical bars on body, like the laces of a corset

Centropyge multicolor Randall & Wass 1974    multi-, many, i.e., multicolored, referring to seven different colors it exhibits in life: white, brown, black, yellow, orange, blue, and violet, more than any other species of the genus

Centropyge multispinis (Playfair 1867)    multi-, many; spinis, spine, referring to 2-4 spines on interoperculum, the “most remarkable characteristic of this fish”

Centropyge nahackyi Kosaki 1989    in honor of “veteran” marine aquarium-fish collector Anthony “Tony” Nahacky; “In his many years of diving in Hawaii and the rest of the Pacific, [he] has made numerous contributions to research on Indo-Pacific reef fishes,” including information on this species

Centropyge narcosis Pyle & Randall 1993    a state of stupor, drowsiness, or unconsciousness, referring to extreme effects of nitrogen narcosis experienced by the senior author and Charles Boyle (see C. boylei) while collecting holotype below 100 m using conventional SCUBA

Centropyge nox (Bleeker 1853)    night, referring to uniformly black color over entire body, head and fins

Centropyge potteri (Jordan & Metz 1912)    in honor of Frederic A. Potter, first director (1904-1940) of the Honolulu (now Waikiki) Aquarium, who presented type to the U.S. National Museum

Centropyge shepardi Randall & Yasuda 1979    in honor of John W. Shepard, University of Guam, who collected C. bispinosa in Guam, thus showing it co-exists with this species, and called the authors’ attention to the difference in the two species’ pectoral-fin ray counts

Centropyge tibicen (Cuvier 1831)    trumpeter (also a piper or flutist), presumably alluding to Japonfche Trompetter, a name given to this angelfish by Dutch naturalist François Valentijn (1666-1727) in 1726; the descriptive significance of “Trompetter” is not evident

Centropyge vrolikii (Bleeker 1853)    in honor of Dutch anatomist-pathologist Willem Vrolik (1801-1863), whose “works on comparative anatomy and developmental history are of great renown”

Centropyge woodheadi Kuiter 1998    in honor of underwater photographer Phil Woodhead (b. 1956), who brought this species to Kuiter’s attention

Subgenus Paracentropyge Burgess 1991    para-, near, proposed as a full genus, “referring to its position near the genus Centropyge

Centropyge boylei Pyle & Randall 1992    in honor of diver and aquarium-fish collector Charles “Chip” J. Boyle, who discovered this angelfish and collected three of the type specimens

Centropyge multifasciata (Smith & Radcliffe 1911)    multi-, many; fasciatus, banded, referring to 10 dark-brown bands across body

Centropyge venusta (Yasuda & Tominaga 1969)    beautiful, described as a “beautiful new species” with yellow and blue colors

Subgenus Xiphypops Jordan & Jordan 1922    xiphos, sword; hypo-, below; ops, eye, referring to two strong spines besides smaller serrae on suborbital bone of C. fisheri

Centropyge acanthops (Norman 1922)    acanthus, spine; ops, eye, presumably referring to strong backwardly directed spine below eye

Centropyge argi Woods & Kanazawa 1951    of Argus Bank, an offshore coral bank about ~25km southwest of Bermuda, type locality

Centropyge aurantonotus Burgess 1974    aurantia, orange; notus, back, referring to golden-yellow head, back and dorsal fin

Centropyge fisheri (Snyder 1904)    in honor of zoologist Walter K. Fisher (1878-1953), Stanford University (California, USA), and member of the 1902 cruise of the U.S. Fish Commission research vessel Albatross, from which type was collected

Centropyge nigriocellus Woods & Schultz 1953    niger, black; ocellus, eye-like spot, referring to black ocellus on base of posterior soft dorsal-fin rays

Centropyge resplendens Lubbock & Sankey 1975    resplendent, referring to its “beautiful” coloration

Incertae sedis

Centropyge abei Allen, Young & Colin 2006    in honor of Yoshitaka Abe (b. 1940), director of Aquamarine Fukushima, a “world class” public aquarium in Japan, who provided the entire budget and material support for the deepwater operations that resulted in the collection of this angelfish; in addition, Abe is “largely responsible for many innovations in aquarium science and design including display of large tunas up to 100kg, jellyfish keeping, and the first public display of hammerhead sharks”                    

Centropyge deborae Shen, Ho & Chang 2012    in honor of Deborah (latinized to Debora) Smith, named at the request of her husband Walt Smith, who collected type series and provided underwater photographs                        

Chaetodontoplus Bleeker 1876    enoplus, armed; Chaetodon, similar to butterflyfishes (Chaetodontidae, in which Bleeker and his contemporaries included both butterflyfishes and angelfishes), referring to prominent spine on posterior edge of gill cover (present in angelfishes but not butterflyfishes)

Chaetodontoplus ballinae Whitley 1959    of Ballina Bar, New South Wales, Australia, type locality

Chaetodontoplus caeruleopunctatus Yasuda & Tominaga 1976    caeruleo-, blue; punctatus, spotted, referring to “minute, but brilliant and numerous” blue spots scattered on sides of living specimens

Chaetodontoplus cephalareticulatus Shen & Lim 1975    cephalo-, head; reticulatus, net-like or netted, allusion not explained, presumably referring to wavy blue stripes (pale in alcohol) on head (if this explanation is correct, then “undulatus” or “vermiculatus” would have been a better adjective)

Chaetodontoplus chrysocephalus (Bleeker 1855)    chrysos, gold; cephalus, head, presumably referring to reddish-orange (actually, yellow-orange) anterior body (i.e., head)

Chaetodontoplus conspicillatus (Waite 1900)    spectacled, presumably referring to blue ring around eye

Chaetodontoplus dimidiatus (Bleeker 1860)    divided in two, referring to adult coloration, light gray on upper half of body, black on lower half

Chaetodontoplus duboulayi (Günther 1867)    in honor of “Duboulay,” who collected type, probably Francis Houssemayne du Boulay (1837-1914), collector, entomologist and natural history artist

Chaetodontoplus melanosoma (Bleeker 1853)    melanos, black; soma, body, referring to “completely black unspotted body” (translation) of adults (contemporary accounts report color ranges from black to gray, with juveniles having a yellow head)

Chaetodontoplus meredithi Kuiter 1990    in honor of John G. Meredith, marine aquarist and dive instructor, who collected type with a handnet                                

Chaetodontoplus mesoleucus (Bloch 1787)    mesos, middle; leucus, white, allusion not explained, perhaps referring to how white anterior half of body intergrades with black posterior half in middle of body

Chaetodontoplus niger Chan 1966    black, referring to “dark to blackish brown” head and body in alcohol, and with a “dark touch of purple” in life                                  

Chaetodontoplus personifer (McCulloch 1914)    persona, mask; fero, to bear, presumably referring to bluish-gray head covered in yellow spots            

Chaetodontoplus poliourus Randall & Rocha 2009    polios, gray; oura, tail, referring to predominantly gray caudal fin, its most distinguishing color feature                    

Chaetodontoplus septentrionalis (Temminck & Schlegel 1844)    northern, allusion not explained, perhaps referring to its slightly more northern distribution (reaching southern Japan) compared to Pomacanthus annularis and P. imperator, its two presumed congeners at the time

Chaetodontoplus vanderloosi Allen & Steene 2004    in honor of Robert (“Rob”) van der Loos (spelled “Vanderloos” by the authors), for his “generosity in providing numerous opportunities to join dive cruises” aboard his live-aboard dive vessel Chertan (Alotau, Papua New Guinea); he also collected type

Genicanthus Swainson 1839    genys, cheek; acanthus, thorn or spine, referring to prominent spine on posterior edge of gill cover (common to all angelfishes) [nests within Centropyge according to some workers]

Genicanthus bellus Randall 1975    beautiful, the “most colorful of all the species of the genus”

Genicanthus caudovittatus (Günther 1860)    caudo-, tail; vittatus, banded, referring to black band on upper and lower caudal-fin lobes of females                                   

Genicanthus lamarck (Lacepède 1802)    in honor of French naturalist Jean-Baptiste Pierre Antoine de Monet, Chevalier de Lamarck (1744-1829), usually known simply as Lamarck [presumably a noun in apposition, without the patronymic “i”]                          

Genicanthus melanospilos (Bleeker 1857)    melanos, black; spilos, mark or spot, referring to black spot mid-ventrally on thorax of males

Genicanthus personatus Randall 1975    masked, referring to dark face on white body of females (males, discovered three years later, have an orange face or “mask”)

Genicanthus semicinctus (Waite 1900)    semi-, half; cinctus, belt or girdle, referring to 8-10 narrow black bars extending to middle of sides of males

Genicanthus semifasciatus (Kamohara 1934)    semi-, half; fasciatus, banded, referring to numerous close-set, wavy bars on upper half of body (not extending to abdomen) of males

Genicanthus spinus Randall 1975    spine, referring to “exceptionally large” anterior preorbital spine

Genicanthus takeuchii Pyle 1997    in honor of diver Hiroshi Takeuchi, who discovered and photographed this angelfish in 1987

Genicanthus watanabei (Yasuda & Tominaga 1970)    in honor of ichthyologist Masao Watanabe, Waseda University (Tokyo, Japan), who first reported this species under the name Holacanthus caudovittatus in 1949                                    

Holacanthus Lacepède 1802   holos, whole or entire; acanthus, thorn or spine, allusion not explained, possibly referring to one or more strong spines on interopercle (compared to none on Pomacanthus, also described by Lacepède)

Subgenus Holacanthus

Holacanthus tricolor (Bloch 1795)    three-colored, referring to adult coloration: yellow anterior body, black posterior body, and (on some specimens) yellow fins edged in red (juveniles are entirely yellow with an ocellated black spot on side that enlarges as it grows)                   

Subgenus Angelichthys Jordan & Evermann 1896    angel, referring to “angelfish,” common name of H. ciliaris; ichthys, fish

Holacanthus africanus Cadenat 1951    African, the first species of the genus reported from the west coast of Africa

Holacanthus bermudensis Goode 1876    ensis, suffix denoting place: proposed as a subspecies of H. ciliaris from Bermuda (also occurs from North Carolina to southern Florida, Bahamas, and Gulf of Mexico)

Holacanthus ciliaris (Linnaeus 1758)    fringed, referring to its ciliate scales (“squamis ciliatis”); name does not mean “with eyelashes,” referring to its “produced” dorsal and anal fins as alleged by Jordan & Evermann (1898) and others

Holacanthus clarionensis Gilbert 1890    ensis, suffix denoting place: Clarion Island, one of three islands (including Socorro and San Benedicto) off western México where this angelfish was “extremely abundant” in 1889

Holacanthus limbaughi Baldwin 1963    in honor of the late Conrad Limbaugh (1925-1960), zoologist, diver and underwater photographer, who was “instrumental” in collecting (with explosives) this angelfish and in charge of the Clipperton Island (type locality in the eastern Pacific) shore parties [Limbaugh died after losing his way while diving in the labyrinth of an underground river in France]

Holacanthus passer Valenciennes 1846    sparrow, allusion not explained nor evident

Pomacanthus Lacepède 1802    poma, lid or cover; acanthus, thorn or spine, referring to prominent spine on posterior edge of gill cover (common to all angelfishes)    

Subgenus Pomacanthus

Pomacanthus arcuatus (Linnaeus 1758)    bowed, referring to 2-3 bow-shaped yellow bands on body of juveniles (absent in adults)

Pomacanthus paru (Bloch 1787)    Portuguese name for this species as reported in Marcgrave’s Historiae naturalis brasiliae (1648)

Pomacanthus zonipectus (Gill 1862)    zona, belt; pectus, breast, referring to dark-brown band “girdling” breast behind ventral and pectoral fins

Subgenus Acanthochaetodon Bleeker 1876    acanthus, thorn or spine; Chaetodon, similar to butterflyfishes (Chaetodontidae, in which Bleeker and his contemporaries included both butterflyfishes and angelfishes), referring to prominent spine on posterior edge of gill cover, common to all angelfishes but which butterflyfishes do not possess

Pomacanthus annularis (Bloch 1787)    ringed, referring to blue ring above gill cover of adults

Pomacanthus chrysurus (Cuvier 1831)    chryso-, gold; oura, tail, referring to yellow caudal fin

Pomacanthus imperator (Bloch 1787)    emporer, based on Keyser van Iapan (Emporer of Japan), Dutch name given to this angelfish by publisher Louis Renard (ca. 1678-1746) in 1719, perhaps alluding to its majestic appearance; Renard called it the “most beautiful fish in the world,” covered with small scales “more brilliant than gold” (translation) [see P. navarchus for another “noble” name coined or reported by Renard]

Pomacanthus rhomboides (Gilchrist & Thompson 1908)    oides, having the form of: a rhombus, allusion not explained, perhaps referring to body shape (with a deep body, unique in the genus)

Subgenus Arusetta Fraser-Brunner 1933    from Aruset el báhr, Arabic name for C. maculosus along the Red Sea of Yemen (type locality)

Pomacanthus asfur (Forsskål 1775)    from Ta-bak el herr, Asfûr, its Arabic name along the Red Sea of Yemen (type locality)

Pomacanthus maculosus (Forsskål 1775)    spotted, referring to blue spots on forehead, nape and operculum of adults (actually blue scales with darker edges) and/or a large gold spot (“macula magna,” but actually a blotch) on middle of body

Pomacanthus semicirculatus (Cuvier 1831)    semi-, half; circulatus, circled, referring to semicircular narrow blue-and-white lines on posterior body of juveniles (absent on adults)

Subgenus Euxiphipops Fraser-Brunner 1934    eu-, well or very; xiphos, sword; hypo-, below; ops, eye, allusion not explained, perhaps referring to strong spines on lower limb of preoperculum of P. xanthometopon

Pomacanthus navarchus (Cuvier 1831)    leader of the ships, based on “Admiral,” Dutch name given to this angelfish by publisher Louis Renard (ca. 1678-1746) in 1719, who said that “more than 30 different kinds” of this species (but perhaps referring to angelfishes and butterflyfishes in general) are known at Ambon, Indonesia, “distinguished by the names royal, imperial, duke, duchess, marquis, count, baron and other titles of nobility” (translation); allusion not explained, possibly (per Mazza 2013, Monaco Nature Encyclopedia) referring to blue stripe crossing operculum, like the blue shoulder strap of an admiral [see P. imperator for another “noble” name coined or reported by Renard]

Pomacanthus sexstriatus (Cuvier 1831)    sex, six; striatus, striped, referring to five broad dark bars on sides and one behind eye of adults (juveniles have white bars)

Pomacanthus xanthometopon (Bleeker 1853)    xanthus, yellow; metopon, forehead, referring to yellow “mask” over eyes of adults

Pygoplites Fraser-Brunner 1933    pyge, rump or rear; [h]oplites, armed (with a weapon), allusion not explained, perhaps referring to “Hind edge of preoperculum dentate or with a few stout spines,” or perhaps a variation on Centropyge, referring to the three anal-fin spines (see above)

Pygoplites diacanthus (Boddaert 1772)    di-, two; acanthus, thorn or spine, referring to its strong opercular spines (one on each side); Boddaert commented, “I have never seen a fish with such sharp thorns” (translation)


Family DREPANEIDAE Sicklefishes

Drepane Cuvier 1831    scythe or sickle, referring to falcate pectoral fins

Drepane africana Osório 1892    African, proposed as a west African (and eastern Atlantic) subspecies of the Indo-West Pacific D. punctata

Drepane longimana (Bloch & Schneider 1801)    longus, long; manus, handed, referring to long, falcate pectoral fins, reaching caudal peduncle

Drepane punctata (Linnaeus 1758)    spotted, referring to brown spots on mid-to-upper body        


Family CHAETODONTIDAE Butterflyfishes
22 genera/subgenera · 138 species/subspecies

Amphichaetodon Burgess 1978    amphi-, on both sides, referring to its two species occurring on opposite sides of the southern Pacific Ocean; Chaetodon, type genus of family

Amphichaetodon howensis (Waite 1903)    ensis, suffix denoting place: Lord Howe Island, type locality (also occurs off southeastern Australia, Norfolk and Kermadec islands, and northern New Zealand)

Amphichaetodon melbae Burgess & Caldwell 1978    matronym not identified but almost certainly in honor of marine mammal biologist Melba C. Caldwell, University of Florida (Gainesville, USA), wife of junior author David K. Caldwell

Chaetodon Linnaeus 1758    chaeto-, bristle; odon, tooth, referring to setiform teeth forming a brush-like bundle

Subgenus Chaetodon

Chaetodon capistratus Linnaeus 1758    bridled or muzzled, allusion not explained, perhaps referring to dark bar running through eye of juveniles and sub-adults

Chaetodon hoefleri Steindachner 1881    in honor of Steindachner’s “dear friend” (translation) W. Höfler (no other information available), who collected or provided type from Senegal, and was one of Steindachner’s best suppliers of fishes from Africa

Chaetodon humeralis Günther 1860    pertaining to the shoulder, allusion not explained, perhaps referring to “brown vertical band from the first, second and third dorsal spines to and below the base of the pectoral, touching the opercular margin”

Chaetodon marleyi Regan 1921    in honor of Natal fisheries officer Harold Walter Bell-Marley (1872-1945), who collected type

Chaetodon ocellatus Bloch 1787    having an eye-like spot, referring to small black spot on soft portion of dorsal fin (although illustrated with a white ring encircling the spot, contemporary photos indicate the spot is not ringed and therefore not a true ocellus)

Chaetodon robustus Günther 1860    full-bodied or strong, allusion not explained, perhaps referring to “much elevated” body and/or “rather strong” fin spines             

Chaetodon striatus Linnaeus 1758    striped, referring to three broad brown bars on sides [provisionally placed in subgenus Chaetodon; probably warrants a subgenus of its own per Smith et al. 2003]

Subgenus Citharoedus Kaup 1860    etymology not explained, presumably cithara-like, allusion not explained, referring to oblique or curved stripes on sides of C. meyeri and/or C. ornatissimus, which can be said to resemble the strings of a cithara (ancient hand-held harp or lute)

Chaetodon meyeri Bloch & Schneider 1801    patronym not identified but type specimen was held in Museo Meyeri Lugduni Batavorum (Meyer’s Museum in Leiden, Netherlands; Meyer may have been Johan Frederik Meijer, a “bookkeeper” at Ambon Island, Indonesia (type locality) and who maintained a small natural-history cabinet in the Netherlands in the 1700s

Chaetodon ornatissimus Cuvier 1831    very ornate, referring to its coloration, including six oblique orange bands on sides, alternating black and gold bars on head, and bluish-white caudal fin with black submarginal band and central black bar                   

Chaetodon reticulatus Cuvier 1831    net-like or netted, referring to black body with lemon-colored scale centers, giving it a reticulated appearance                                

Subgenus Corallochaetodon Burgess 1978    corallo-, coral, a subgenus of Chaetodon known for feeding on live coral polyps (Warren E. Burgess, pers. comm.)                           

Chaetodon austriacus Rüppell 1836    etymology not explained; usually austriacus means Austrian but that does not make sense for a Red Sea fish; perhaps name is a derivative of auster or australis, meaning south, possibly referring to relative position of type locality (Jeddah, Saudi Arabia) to the Sinai, where many butterflyfishes had been found washed ashore (Ronald Fricke, pers. comm.)

Chaetodon lunulatus Quoy & Gaimard 1825    somewhat moon-shaped, presumably referring to its suborbicular body shape

Chaetodon melapterus Guichenot 1863    melas, black; pterus, fin, referring to black dorsal, caudal and anal fins

Chaetodon trifasciatus Park 1797    tri-, three; fasciatus, banded, referring to three black bars on head: on lower jaw, through eye, and behind eye [not to be confused with C. trifascialis]

Subgenus Discochaetodon Nalbant 1971    disco-, disk, subgenus of Chaetodon differing from the “closely related” subgenus Citharoedus by its “almost circular body”                                    

Chaetodon aureofasciatus Macleay 1878    aureus, golden; fasciatus, banded, referring to dark-edged golden-orange bar through eye, and another along posterior edge of operculum, through base of pectoral fins to the belly

Chaetodon octofasciatus Bloch 1787    octo-, eight; fasciatus, banded, referring to eight black vertical bars on body, first one through eye, last one along rear edge of dorsal fin

Chaetodon rainfordi McCulloch 1923    in honor of Edward Henry Rainford (1853-1938), amateur naturalist and viticulturist, Queensland Agricultural Department, for whom the Australian Museum is “indebted for many remarkable fishes not previously recognised from Australian waters,” including type of this species

Chaetodon tricinctus Waite 1901    tri-, three; cinctus, belt or girdle, referring to three vertical black bands crossing body

Subgenus Exornator Nalbant 1971   adorner or embellisher, allusion not explained, perhaps referring to color pattern of body “based on vertical or oblique rows of dusky dots or crossbands”

Chaetodon argentatus Smith & Radcliffe 1911    silvery, referring to color in alcohol, “Silvery, tinged with a very pale lemon yellow”

Chaetodon assarius Waite 1905    roasted (i.e., browned), allusion not explained, perhaps referring to silvery-tan body with five dark, broken vertical lines on upper sides, reminiscent of the sear marks on grilled meat

Chaetodon blackburnii Desjardins 1836    in honor of Edward Berens Blackburn (1786-1839), English jurist who served as a judge on Mauritius (type locality), then a British Crown Colony; Blackburn was honored as the “learned and virtuous magistrate who presided over” (translation) the Société d’Histoire Naturelle de l’Ile Maurice

Chaetodon citrinellus Cuvier 1831    lemon-colored, referring to white-to-yellow ground color

Chaetodon daedalma Jordan & Fowler 1902    a piece of art embroidery, referring to scales, their centers pale yellow and edges broadly margined with blackish brown, “forming a beautiful reticulated or netted pattern”

Chaetodon dolosus Ahl 1923    Latin for deceitful or “deceptively given” (translation), referring to its misidentification as C. miliaris by Sauvage in 1891

Chaetodon fremblii Bennett 1828    in honor of Lieut. John Frembly, British Royal Navy, marine surveyor and geologist, who presented to the Zoological Society of London an “interesting” collection of fishes from the Sandwich Islands (Hawai‘i), including type of this one

Chaetodon guentheri Ahl 1923    in honor of ichthyologist-herpetologist Albert Günther (1830-1914), who reported this species as C. miliaris in 1871

Chaetodon guttatissimus Bennett 1833    full of dots, referring to close-set gray spots or speckles on sides

Chaetodon litus Randall & Caldwell 1973    plain or simple, referring to its plain color, “very much in contrast to the striking color patterns of most butterflyfishes”

Chaetodon madagaskariensis Ahl 1923    ensis, suffix denoting place: based on description of C. miliaris by Sauvage (1891) allegedly from Madagascar but actually from Mauritius [many contemporary authors have changed spelling to madagascariensis; original spelling is preferred but prevailing usage may apply]                                 

Chaetodon mertensii Cuvier 1831    in honor of German naturalist and explorer Karl Heinrich Mertens (1796-1830); Cuvier described the species, type locality unknown, from an illustration that Mertens showed him (whether Mertens was himself the artist is not clear; see also Salmoniformes: Thymallus mertensii)

Chaetodon miliaris Quoy & Gaimard 1825    arius, pertaining to: mille, thousand, allusion not explained, probably referring to body “covered with a crowd of small black dots forming vertical or horizontal lines” (translation)

Chaetodon multicinctus Garrett 1863    multi-, many; cinctus, belt or zone, presumably referring to five vertical, yellowish-brown, diffuse stripes on sides

Chaetodon paucifasciatus Ahl 1923    pauci-, few; fasciatus, banded, proposed as a subspecies of C. chrysurus (=madagaskariensis) with fewer bands on sides (6 vs. 7-8)

Chaetodon pelewensis Kner 1868    ensis, suffix denoting place: erroneously named for Pelew-Inseln, German spelling of Palau Islands, western Pacific, type locality (does not occur there; actual type locality is probably Society Islands in the South Pacific per Eschmeyer’s Catalog of Fishes) [treated as a junior synonym of C. punctatofasciatus by some workers]

Chaetodon punctatofasciatus Cuvier 1831    punctatus, spotted; fasciatus, banded, referring to ~7 vertical dark bars on upper body and rows of dark spots below

Chaetodon quadrimaculatus Gray 1831    quadri-, four; maculatus, spotted, presumably referring to white spots on middle of and near tail (two on each side, for a total of four) [originally spelled “4-maculatus” but that was emended since zoological names cannot include numerals]

Chaetodon sanctaehelenae Günther 1868    of Saint Helena Island, South Atlantic, type locality (also occurs at Ascension Island and in the eastern Atlantic straying to Canary Islands and Liberia)

Chaetodon sedentarius Poey 1860    sedentary, i.e., prone to stay in place, allusion not explained but probably named to distinguish this Atlantic species previously identified by Guichenot (1853) as the otherwise Indo-West Pacific C. vagabundus, whose name means “wandering”

Chaetodon smithi Randall 1975    in honor of ichthyologist C. Lavett Smith, Jr. (1927-2015), American Museum of Natural History, who collected the first specimens and realized they were an undescribed species

Chaetodon xanthurus Bleeker 1857    xanthos, yellow; oura, tail, referring to orange outer part of caudal fin

Subgenus Gonochaetodon Bleeker 1876    gonio-, angle, probably referring to rhomboidal or roughly triangular body shape of C. triangulum; Chaetodon, type genus of the family          

Chaetodon baronessa Cuvier 1829    derived from douwing-baroness, name for this species used by Dutch sea captain Willem de Vlamingh (1640-ca. 1698), allusion not explained but similar to douwing-princesse, name given to this species by publisher Louis Renard (ca. 1678-1746) in 1719, who said it was “delightful to look at” (translation), representing one of several “noble” names reported or coined by Renard (see Pomacanthus navarchus above in Pomacanthidae); douwing (also spelled doewing) is a local Malay name applied to several angelfish and butterflyfish species             

Chaetodon larvatus Cuvier 1831    masked, referring to brick-red head and snout

Chaetodon triangulum Cuvier 1831    triangle, referring to its roughly triangular body shape

Subgenus Lepidochaetodon Bleeker 1876    lepido-, scale, allusion not explained, perhaps referring to nearly angular or rhomboidal scales compared to related taxa (Chaetodontops, Citharoedus, Rabdophorus) Bleeker described as having rounded scales; Chaetodon, type genus of the family

Chaetodon interruptus Ahl 1923    interrupted, proposed as a subspecies of C. unimaculatus, allusion not explained, perhaps referring to black band through eye ending at edge of preopercle instead of extending onto chest as in C. unimaculatus

Chaetodon kleinii Bloch 1790    in honor of Jacob Theodor Klein (1685-1759), German jurist, historian, botanist, zoologist and mathematician, who wrote a 5-volume history of fishes (1740-1749); Klein’s account and illustration of this species in vol. 4 (1744) was Bloch’s introduction to it

Chaetodon trichrous Günther 1874    tri-, three; chroa, color of body or skin, allusion not explained, illustrated as having a white head, followed by black ocular band, then three different colors for remainder of body, white, black and yellow (modern photographs show only a yellow caudal fin)

Chaetodon unimaculatus Bloch 1787   uni-, one; maculatus, spotted, referring to large black spot on upper sides

Subgenus Megaprotodon Guichenot 1848    mega-, large; protos, in front of or forward; odon, tooth, referring to a “group of teeth longer than the others, as if ended in hooks, and placed towards the end [front end?] of the lower jaw” (translation) of C. bifascialis (=trifascialis)

Chaetodon trifascialis Quoy & Gaimard 1825    tri-, three; fascialis, banded, referring to three vertical black bands on juveniles: (1) crossing cheek and passing over eye, (2) posterior body from dorsal fin though anal (juveniles only), (3) submarginally on caudal fin (adults lose bands 2 and 3) [not to be confused with C. trifasciatus]

Subgenus Rabdophorus Swainson 1839    rhabdos, rod; ophorus, bearer, referring to dorsal fin of C. ephippium “prolonged into a whip-like filament”

Chaetodon adiergastos Seale 1910    Greek for unfinished, allusion not explained but described as related to C. flavirostris, “but lack[ing] the wide, dusky band from dorsal to anal which characterizes that species”

Chaetodon auriga Forsskål 1775    coachman, referring to trailing filament of soft dorsal fin of adults, like a coachman’s whip

Chaetodon auripes Jordan & Snyder 1901    aureus, gold; pes, foot, allusion not explained, perhaps referring to golden-yellow pelvic (ventral) fins

Chaetodon collare Bloch 1787    collar, referring to white bar behind eye and narrow white line behind mouth, both wrapping around head like a collar                           

Chaetodon decussatus Cuvier 1829    divided crosswise in the form of an X, referring to oblique lines set at right angles to each other forming a chevron-like pattern on the sides

Chaetodon dialeucos Salm & Mee 1989    dia-, through or between; leucos, white, i.e., intermixed with white, referring to triangular white band between black band through eye and gray-brown body

Chaetodon ephippium Cuvier 1831    saddle, presumably referring to large, black saddle-shaped patch on rear upper corner of body

Chaetodon falcula Bloch 1795    diminutive of falx or falcis, i.e., small sickle or scythe, referring to pair of prominent wedge-shaped black bars on upper sides, which, in Bloch’s illustration, are shaped like sickles (but not shaped that way as shown in contemporary photos)

Chaetodon fasciatus Forsskål 1775    banded, allusion not explained, probably referring to black patch surrounding eye and connecting across forehead and/or diagonal black bands on sides

Chaetodon flavirostris Günther 1874    flavus, yellow; rostris, snout, referring to yellow-orange band encircling snout

Chaetodon gardineri Norman 1939    in honor of British zoologist John Stanley Gardiner (1872-1946), Secretary to the Committee of the John Murray Expedition to the Indian Ocean, during which type was collected

Chaetodon leucopleura Playfair 1867    leukos, white; pleura, side, presumably referring to “white or yellowish on the middle of the sides”

Chaetodon lineolatus Cuvier 1831    lined, referring to series of vertical black lines on sides

Chaetodon lunula (Lacepède 1802)    little moon, presumably referring to black crescent-shaped spot over each eye

Chaetodon melannotus Bloch & Schneider 1801    melanos, black; notus, black, presumably referring to black dorsal portion of body (except for two white patches), usually only seen at night or when fish is frightened [spelling emended to the orthographically correct “melanotus” by some authors but original spelling stands]

Chaetodon mesoleucos Forsskål 1775    meso-, middle; leucos, white, although this translation does not quite match the fish’s color pattern, which is silvery white on head and front part of body and mostly grayish-brown on remainder of body                          

Chaetodon nigropunctatus Sauvage 1880    nigro-, black; punctatus, spotted, referring to black scale centers, which give it a spotted appearance

Chaetodon ocellicaudus Cuvier 1831    ocellus, eye-like spot; caudus, tail, referring to black ocellus on caudal peduncle

Chaetodon oxycephalus Bleeker 1853    oxy, sharp; cephalus, head, referring to “pointed profile” (translation) of its projecting snout

Chaetodon pictus Forsskål 1775    painted, allusion not explained, perhaps referring to two series of narrow, oblique violet lines set at right angles to each other across a whitish body

Chaetodon rafflesii Anonymous [Bennett] 1830    in honor of British statesman Thomas Stamford Raffles (1781-1826), Lieutenant-Governor of the Dutch East Indies (1811-1816), founder of modern Singapore and first President of the Zoological Society of London; he employed zoologists and botanists to collect specimens, paying them out of his own pocket [description appeared in an appendix to a memoir written by his widow]

Chaetodon selene Bleeker 1853    moon, allusion not explained, perhaps referring to crescent-shaped black band running along dorsal-fin base, through caudal peduncle, to middle anal-fin base

Chaetodon semeion Bleeker 1855    banner, presumably referring to filament trailing from soft portion of dorsal fin

Chaetodon semilarvatus Cuvier 1831    semi-, half; larvatus, masked, referring to dark bluish-gray patch behind eye, covering parts of operculum and cheeks but not front of face

Chaetodon ulietensis Cuvier 1831    ensis, suffix denoting place: Uliétéa (now known as Raiatea) Island, French Polynesia, South Pacific, type locality (but widely occurs in central and western Pacific)

Chaetodon vagabundus Linnaeus 1758    wandering, allusion not explained nor evident (although it is very widespread throughout the Indo-West and central Pacific)

Chaetodon wiebeli Kaup 1863    in honor of Kaup’s “dear friend” (translation) Karl Maximilian Wiebel (1808-1888), German naturalist and a founding member of the Zoological Museum in Hamburg (which helped Kaup acquire a specimen)                      

Chaetodon xanthocephalus Bennett 1833    xanthus, yellow; cephalus, head, referring to yellow-orange head (and breast)

Subgenus Roaops Maugé & Bauchot 1984    ops, appearance, referring to similarity to (and previous placement within) the subgenus Roa (now a full genus)

Chaetodon burgessi Allen & Starck 1973    in honor of ichthyologist Warren E. Burgess, who, while preparing his worldwide revision of the family (published 1978), brought the authors’ attention to the close affinity of this species to C. mitratus                           

Chaetodon declivis declivis Randall 1975    Latin for sloping, referring to diagonally demarcated orange patch on upper rear portion of body and fins                            

Chaetodon declivis wilderi Pyle 1988    in honor of David Wilder, an aquarium-fish collector at Christmas Island (type locality), “one of the first to observe and collect” this subspecies

Chaetodon flavocoronatus Myers 1980    flavus, yellow; coronatus, crowned, referring to crown-like yellow band on nape

Chaetodon mitratus Günther 1860    wearing a hand band, presumably referring to ocular band, which “crosses the head above the orbits and does not extend to the interoperculum”

Chaetodon nippon Steindachner & Döderlein 1883    Japanese name for Japan (described from Tokyo and common in south Japan but occurs along western Pacific from northern Philippines to South Korea)

Chaetodon tinkeri Schultz 1951    in honor of Spencer Wilkie Tinker (1909-1999), Director, Waikiki Aquarium (Honolulu), who provided type                     

Subgenus Tetrachaetodon Weber & de Beaufort 1936    tetra-, four, i.e., a subgenus of Chaetodon proposed for C. plebeius, then distinguished by having four (sometimes five) anal-fin spines

Chaetodon andamanensis Kuiter & Debelius 1999    ensis, suffix denoting place: Andaman Sea, eastern Indian Ocean, where type locality (Great Nicobar Island) is situated

Chaetodon bennetti Cuvier 1831    in honor of zoologist Edward Turner Bennett (1797-1836), Zoological Society of London, who showed type specimen to Cuvier

Chaetodon plebeius Cuvier 1831    common, described but not named by Gmelin 1789, who said it was very common (“numerosissima”)

Chaetodon speculum Cuvier 1831    mirror, manuscript name coined by Kuhl & van Hasselt, allusion not explained nor evident, perhaps referring to “very brilliant golden yellow” (translation) body color or oval black spot on middle of upper sides (oval being a common shape for decorative wall mirrors)

Chaetodon zanzibarensis Playfair 1867    ensis, suffix denoting place: Zanzibar, Tanzania, western Indian Ocean, type locality (occurs from KwaZulu-Natal, South Africa east to Chagos Archipelago)

Chelmon Cloquet 1817    based on “Chelmons” of Cuvier 1816, from name of an unknown fish used by Hesychius of Alexandria, a Greek grammarian (probably 5th or 6th century AD), which Cuvier applied to C. rostratus

Chelmon marginalis Richardson 1842    edged or bordered, referring to submarginal dark bands (orange and blue in life) around soft portions of dorsal and anal fins (absent in the similar C. rostratus)

Chelmon muelleri Klunzinger 1879    in honor of physician, geographer and botanist Ferdinand von Mueller (1825-1896), who donated his extensive collection of Australian plant and animal specimens, including type of this species, to what is now the Staatliches Museum für Naturkunde in Stuttgart (Baden-Württemberg, Germany)

Chelmon rostratus (Linnaeus 1758)    beaked, referring to elongated snout (an adaptation for feeding on benthic invertebrates in small crevices)                          

Chelmonops Bleeker 1876    ops, appearance, referring to similarity to (and previous placement of C. truncatus in) Chelmon

Chelmonops curiosus Kuiter 1986    curious, referring to its “inquisitive nature towards divers, often looking into their mask or camera lens when confronted”                 

Chelmonops truncatus (Kner 1859)    truncate, referring to its elongated snout (for a Chaetodon, original genus) but shorter than those of the genus Chelmon

Coradion Kaup 1860    kore, pupil of the eye; adion, placed sideways (on the flank), allusion not explained, perhaps referring to ocellus (eye-spot) at base of soft dorsal fin of C. chryzozonus

Coradion altivelis McCulloch 1916    altus, high; velum, sail, allusion not explained but clearly referring to elevated soft dorsal and anal fins

Coradion calendula Matsunuma, Motomura & Seah 2023    Calendula, generic name of plants in the daisy family Asteraceae (often known as marigolds), referring to the characteristic orange band on the caudal peduncle

Coradion chrysozonus (Cuvier 1831)    chrysos, gold; zonus, belt or girdle, referring to three brownish-orange bars on sides

Coradion melanopus (Cuvier 1831)    melanos, black; pous, feet, referring to dark pelvic (ventral) fins

Forcipiger Jordan & McGregor 1898    forceps, tweezers; –iger, to bear, referring to long and slender snout, with short, forcep-like jaws at the end

Forcipiger cyrano Randall 1961    etymology not explained but almost certainly referring to Cyrano de Bergerac, main character in an 1897 comedy by Edmond Rostand, characterized by his large nose, alluding to its long snout, even longer than the “absurdly” long and slender snout of F. longirostris

Forcipiger flavissimus Jordan & McGregor 1898    very yellow, referring to bright yellow body color (upper half of head and nape are black)

Forcipiger longirostris (Broussonet 1782)    longus, long; rostris, snout, referring to long and slender snout, with which it packs small invertebrates from within corals or from reef crevices

Forcipiger wanai Allen, Erdmann & Jones Sbrocco 2012    name for this butterflyfish used by local inhabitants (Wandammen language group) in the southwestern part of Cenderawasih Bay, West Papua Province, Indonesia, type locality

Hemitaurichthys Bleeker 1876    hemi-, partial, presumably referring to similarity to and/or relationship with Taurichthys (=Heniochus) but with more lateral-line scales (taurus, bull; ichthys, fish, originally referring to horn-like bony protuberances on head of Heniochus varius)

Hemitaurichthys multispinosus Randall 1975    multi-, many; spinosus, spiny, referring to higher number of dorsal- and anal-fin spines compared to congeners                    

Hemitaurichthys polylepis (Bleeker 1857)    poly, many; lepis, scale, referring to numerous lateral-line scales (68-74), then considered the most of any of its presumed congeners in Chaetodon (original genus)

Hemitaurichthys thompsoni Fowler 1923    in honor of John W. Thompson, artist and modeler at the Bishop Museum (Honolulu), who obtained type at a fish market

Hemitaurichthys zoster (Bennett 1831)    belt or girdle, presumably referring to black body crossed by a broad, white trapezoid band

Heniochus Cuvier 1816    driver (of a horse-drawn carriage), referring to filamentous fourth dorsal-fin spine of H. macrolepidotus (=acuminatus), “similar to a kind of whip” (translated from Cuvier 1829)

Heniochus acuminatus (Linnaeus 1758)    pointed or sharpened, allusion not explained, perhaps referring to elongate fourth dorsal-fin spine, filamentous and often longer than body length

Heniochus chrysostomus Cuvier 1831    chrysos, gold; stomus, mouth, referring to yellow snout

Heniochus diphreutes Jordan 1903    charioteer, allusion not explained but almost certainly referring (like the name of the genus) to elongate dorsal-fin filament similar to a horseman’s whip

Heniochus intermedius Steindachner 1893    intermediate in shape and color between H. macrolepidotus (=acuminatus) and H. chrysostomus

Heniochus monoceros Cuvier 1831    mono-, one; ceros, horn, referring to prominent bony protuberance on head

Heniochus pleurotaenia Ahl 1923     pleuro-, side; taenia, band, referring to two broad black bands on lower sides

Heniochus singularius Smith & Radcliffe 1911    unique, single, remarkable or unusual, allusion not explained nor evident

Heniochus varius (Cuvier 1829)    etymology not explained, perhaps meaning different, referring to unusual appearance of its elevated head, two horn-like bony protuberances at level of eyes, and another protuberance on forehead (varius could also mean varied or variegated, referring to its color pattern, but its largely brown and black coloration with two white bands does not seem to match that adjective)

Johnrandallia Nalbant 1974    ia, belonging to: John E. Randall (1924-2020), Bishop Museum (Honolulu), in appreciation of his work on reef fishes 

Johnrandallia nigrirostris (Gill 1862)    nigri-, black; rostris, snout, referring to blackish band on snout

Parachaetodon Bleeker 1874    para-, referring to similarity to Chaetodon and/or previous placement of P. oligacanthus (=ocellatus) in that genus

Parachaetodon ocellatus (Cuvier 1831)    having an eye-like spot, presumably referring to brown ring bordered with white at tail of juveniles (a black oval-shaped mark with a silver anterior edge in adults) [treated as a junior synonym of Chaetodon oligacanthus by some workers]

Prognathodes Gill 1862    oides, having the form of: pro-, forward; gnathus jaw, presumably referring to produced snout of P. pelta (=aculeatus)

Prognathodes aculeatus (Poey 1860)    spined, presumably referring to long and very strong dorsal-fin spines

Prognathodes aya (Jordan 1886)    named for the Red Snapper, Lutjanus aya (=campechanus), from whose stomach type specimen was “spewed up”                 

Prognathodes basabei Pyle & Kosaki 2016    in honor of Peter K. Basabe, “long-time diver, aquarium fish collector and resident of Kona, Hawai‘i, both for his role in the collection of the first specimen of this new species in 1998 and more generally for his extensive contributions and assistance to many researchers (especially the authors) in the ichthyological community”

Prognathodes brasiliensis Burgess 2001    ensis, suffix denoting place: Brazil, where type locality (Cabo Frio, near Rio de Janeiro) is situated, and country that “probably defines its range”

Prognathodes carlhubbsi Nalbant 1995    in memory of ichthyologist Carl L. Hubbs (1894-1979), who was planning to describe this species before he died; he also published a study of this genus in 1963 and co-described P. falcifer in 1958                      

Prognathodes dichrous (Günther 1869)    di-, two; chroa, color of body or skin, i.e., bicolored, chocolate brown with conspicuous rectangular white patch on upper rear corner of body and adjacent dorsal fin

Prognathodes falcifer (Hubbs & Rechnitzer 1958)    falcis, sickle or scythe; fero, to bear, referring to scythe-shaped black mark from gill cover to anal fin

Prognathodes geminus Copus, Pyle, Greene & Randall 2019    twin, referring to its similarity in color to P. basabei of the Hawaiian Islands

Prognathodes guezei (Maugé & Bauchot 1976)    in honor of marine biologist Paul Guézé, who collected holotype and helped collect paratypes                              

Prognathodes guyanensis (Durand 1960)    ensis, suffix denoting place: French Guiana, type locality (occurs in western Atlantic from Bahamas south to Rio Grande do Sul, Brazil)

Prognathodes guyotensis (Yamamoto & Tameka 1982)    ensis, suffix denoting place: referring to its “peculiar unexpected distribution on the top of a guyot of 330m deep, which is by far the deepest record” of a chaetodontid fish                                  

Prognathodes marcellae (Poll 1950)    in honor of Poll’s wife Marcelle Aen Poll (née den Boom), his “generous and dedicated collaborator” (translation)                         

Prognathodes obliquus (Lubbock & Edwards 1980)    slanting, referring to white patch on upper rear corner of body, running slantwise from base of exposed portion of sixth dorsal-fin spine to base of last anal-fin ray

Roa Jordan 1923    Polynesian (New Zealand, Tonga, Tahiti) word meaning high or long, referring to type specimen of R. excelsa, a deepwater fish killed and brought to the surface by a lava stream from Mauna Loa volcano in Hawai‘i (loa means high or long in Hawaiian) [replacement name for Loa Jordan 1921, preoccupied by Loa Stiles 1905 in nematodes]

Roa australis Kuiter 2004    southern, referring to distribution in the Southern Hemisphere (northwestern Australia and Arafura Sea)

Roa excelsa (Jordan 1921)    Latin for high or lofty, referring to type specimen, a deepwater fish killed and brought to the surface by a lava stream from Mauna Loa volcano in Hawai‘i (loa means high or long in Hawaiian)

Roa haraguchiae Uejo, Senou & Motomura 2020    in honor of Mrs. Yuriko Haraguchi, a volunteer at the Kagoshima University Museum for over 15 years, who has “kindly supported” the authors’ ichthyological research and fish-collection management

Roa jayakari (Norman 1939)    in honor of Atmaram Sadashiva Grandin Jayakar (1844-1911), surgeon, Indian Medical Service, and collector of natural history, who sent four specimens to the British Museum

Roa modesta (Temminck & Schlegel 1844)    modest or unassuming: “The colors of this fish are not very bright” (translation)

Roa rumsfeldi Rocha, Pinheiro, Wandell, Rocha & Shepherd 2017    in honor of Donald Rumsfeld (1932-2021), Secretary of Defense under U.S. presidents Gerald Ford and George W. Bush and a key figure in the planning of the Afghanistan and Iraq wars, who immortalized the quote, “there are known knowns; there are things we know we know. We also know there are known unknowns; that is to say we know there are some things we do not know. But there are also unknown unknowns – the ones we don’t know we don’t know”; according to the authors, Rumfeld’s quote about the “uncertainties of war” applies perfectly to the taxonomy of MCE (mesophotic coral ecosystem) species: “We only realized this species was new after we took a good look at it here at the aquarium in San Francisco,” the authors wrote, “so we think it’s a perfect example of an unknown unknown.”

Roa semilunaris Matsunuma & Motomura 2022    semi-, from semis, half; lunaris, of the moon, referring to halfmoon-shaped ocellated blotch on dorsal fin

Roa uejoi Matsunuma & Motomura 2022    in honor of Takuya Uejo, Kagoshima University (Japan), who first recognized the uniqueness of this species while studying the taxonomy of the genus