Family CYPRINIDAE: Subfamily SCHIZOPYGOPSINAE Mirza 1991 (Mountain Barbels)

Updated 2 May 2023
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Diptychus Steindachner 1866 di- (Gr. prefix), from dýo (δύο), two; ptychus, from ptychós (Gr. πτυχός), genitive of ptýx (πτύξ), layer or fold, referring to lower jaw of D. maculatus, with a cutting anterior edge covered with a horny sheath, behind which is a distinct, uninterrupted labial fold, continuous with upper lip

Diptychus maculatus Steindachner 1866 Latin for spotted, referring to two rows of dark speckles on abdomen and lower portion of head

Diptychus sewerzowi Kessler 1872 in honor of Russian zoologist Nikolai A. Severtzov (also spelled Severzov and Sewerzoff, 1827–1885), who collected holotype

Gymnodiptychus Herzenstein 1892 gymnos, bare or naked, referring to absence of scales except on lateral line; Diptychus, genus in which type species, G. dybowskii, had originally been assigned

Gymnodiptychus dybowskii (Kessler 1874) in honor of Polish biologist Benedykt Dybowski (1833–1930), “indefatigable researcher of the fauna of East Siberia” (translation)

Gymnodiptychus integrigymnatus Mo 1989 integer (L.), whole or entire; gymnatus, presumably from gymnós (Gr. γυμνός), bare or naked, referring to “complete” absence of scales (except for anal scales and pelvic axillary scale on body) [authorship credited to Huang but, per Eschmeyer’s Catalog of Fishes, Mo wrote the description]

Gymnodiptychus pachycheilus Herzenstein 1892 pachýs (Gr. παχύς), thick; cheilus, from cheī́los (Gr. χεῖλος), lip, probably referring to thick and fleshy lower lip

Gymnodiptychus pachycheilus weiheensis Wang & Song 1985ensis, Latin suffix denoting place: Weihe River system, Gansu Province, China, type locality

Oxygymnocypris Tsao 1964 oxýs (Gr. ὀξύς), sharp or pointed, proposed as a subgenus of Gymnocypris (= Schizopygopsis)with a conical, pointed snout

Oxygymnocypris stewartii (Lloyd 1908) in honor of Capt. Francis Hugh Stewart (1879–1951), surgeon-naturalist, Indian Medical Service, who collected holotype

Ptychobarbus Steindachner 1866 ptychós (Gr. πτυχός), genitive of ptýx (πτύξ), layer or fold, presumably referring to broad, uninterrupted labial fold on P. conirostris; barbus (L.), beard, probably referring to pair of maxillary barbels that extend to hind end of preopercle in adults, and/or to Barbus (Barbinae), then a catch-all genus for many African, Asian and European cyprinids

Ptychobarbus chungtienensis (Tsao 1964) -ensis, Latin suffix denoting place: Zhongdian (phonetically spelled Chungtien) County, Yunnan Province, China, type locality

Ptychobarbus chungtienensis gezaensis (Huang & Chen 1986)ensis, Latin suffix denoting place: Geza, Zhongdian County, Yunnan Province, China, type locality [treated as a full species or a synonym of P. chungtienensis depending on the reference; provisionally included here]

Ptychobarbus conirostris Steindachner 1866 conus, from kṓnos (Gr. κῶνος), cone; rostris, Neo-Latin scientific adjective of rostrum (L.), snout, referring to its conical snout

Ptychobarbus dipogon (Regan 1905) di- (Gr. prefix), from dýo (δύο), two; pṓgōn (Gr. πώγων), beard, referring to one pair of barbels, compared with two pair among its presumed congeners in Schizothorax (Schizothoracinae)

Ptychobarbus kaznakovi Nikolskii 1903 in honor of Russian zoologist Alexander N. Kaznakov (1872–1933), an associate of Pyotr K. Kozlov’s Imperial Russian Geographical Society expedition to Mongolia and Tibet (1899–1901), which yielded a rich natural history collection, including holotype of this species

Ptychobarbus leptosomus Zhang, Zhao & Niu 2019 slender-bodied, from leptós (Gr. λεπτός), thin, and sṓma (Gr. σῶμα), body, referring to its overall slender appearance

Schizopygopsis Steindachner 1866 ópsis (Gr. ὄψις), appearance: similar to Schizopyge (Schizothoracinae), particularly the membranous fold in front of the anal fin

Schizopygopsis anteroventris Wu & Tsao 1989 antero- (L.), opposite; ventris (L.), genitive of venter, belly, probably referring to how ventral (pelvic) fins are directly opposite base of first dorsal-fin ray

Schizopygopsis bangongensis Wu & Zhu 1979ensis, Latin suffix denoting place: Ban-Gong Lake, Tibet, China, type locality

Schizopygopsis chengi (Fang 1936) in honor of botanist Cheng Wan-Chun (1908–1987), Herbarium of the Biological Laboratory of the Science Society of China, who collected holotype

Schizopygopsis chengi baoxingensis Fu, DIng & Ye 1994ensis, Latin suffix denoting place: Baoxing County, Sichuan, China, type locality [proposed as a subspecies of S. malacanthus and treated as such by some workers]

Schizopygopsis chilianensis (Li & Chang 1974)ensis, Latin suffix denoting place: Qilian Range, northeast ridge of Tibetan Plateau, where it occurs (“chilian” is the approximate phonetic spelling of “Qilian”)

Schizopygopsis chui (Tchang, Yueh & Hwang 1964) patronym not identified, possibly in honor of Yuan-Ting Chu (1896–1986), director, Shanghai Fisheries Institute, who published many papers on Chinese fishes in the 1930s

Schizopygopsis dobula (Günther 1868) Latinization of Döbel, German common name for Squalius cephalus (Leuciscidae), which it presumably resembles

Schizopygopsis eckloni (Herzenstein 1891) in honor of F. L. Ecklon, one of Nikolai Przhevalsky’s (see S. przewalskii) assistants during his second trip to Tibet, whose services Przhevalsky said were “invaluable” (translation)

Schizopygopsis eckloni scoliostomus (Wu & Chen 1979) curve-mouthed, from skoliós (Gr. σκολιός), curved or bent, and stóma (Gr. στόμα), referring to “deeply arched” mouth cleft [treated as a full species by some workers]

Schizopygopsis extrema (Herzenstein 1891) Latin for extreme, referring to the distinctiveness of its pharyngeal teeth, described as an “extreme form” (translation) of the teeth found in Gymnocypris (=Schizopygopsis)

Schizopygopsis firmispinata (Wu & Wu 1988) firmus (L.), strong; spinata (L.), spined or spiny, referring to stronger third simple ray of dorsal fin compared with weaker dorsal spine of G. potanini

Schizopygopsis himalayensis Tsao 1974ensis, Latin suffix denoting place: referring to its occurrence near Zhumulangma Mountain (Mt. Everest) in the Himalayas of Tibet

Schizopygopsis kessleri Herzenstein 1891 in memory or Russian ichthyologist Karl Federovich Kessler (1815–1881)

Schizopygopsis kialingensis Tsao & Tun 1962ensis, Latin suffix denoting place: Jialing River, Gansu Province, China, type locality

Schizopygopsis labiosa (Herzenstein 1891) Latin for large-lipped, referring to its “rather fleshy lips” (translation)

Schizopygopsis longimandibularis (Tsao, Chen, Wu & Zhu 1992) longus (L.), long; mandibularis (L.), of or pertaining to a jaw, presumably referring to its longer mouth (reflected in smaller mouth width-to-length ratio) compared with S. chui

Schizopygopsis malacanthus Herzenstein 1891 malakós (Gr. μαλακός), soft; acanthus (L.), from ákantha (Gr. ἄκανθα), thorn, presumably referring to the “very little thickened” dorsal spine on larger individuals in which serrations are “more or less atrophied” (translations)

Schizopygopsis maphamyumensis Wu & Zhu 1979ensis, Latin suffix denoting place: Mapham Yu Tso (Lake Manasarovar), Tibet, where it is endemic

Schizopygopsis microcephalus Herzenstein 1891 small-headed, from mikrós (Gr. μικρός), small, and kephalḗ (Gr. κεφαλή), head, referring to its smaller head compared with S. guentheri (=pylzovi) [reported to occur at 5350 meters, making it the highest-living fish in the world]

Schizopygopsis namensis Wu & Ren 1982ensis, Latin suffix denoting place: Nam Cuo Lake, northeastern Tibet, type locality

Schizopygopsis pengquensis (Tang, Feng, Wanghe, Li & Zhao 2016)ensis, Latin suffix denoting place: Pengqu River, Nielamu and Dingri counties, Tibet, where it appears to be endemic

Schizopygopsis potanini (Herzenstein 1891) in honor of Grigory Nikolayaevich Potanin (1835–1920), Russian botanist, ethnologist and explorer in Central Asia

Schizopygopsis przewalskii Kessler 1876 in honor of Nikolai Przhevalsky (also spelled Przewalski and Prjevalsky, 1839–1888), Russian Cossack geographer, explorer and naturalist, who collected holotype and in whose book about Mongolia Kessler’s description appeared

Schizopygopsis przewalskii ganzihonensis Zhu & Wu 1975ensis, Latin suffix denoting place: Ganzi River, Qinghai, China, where it appears to be endemic

Schizopygopsis pylzovi Kessler 1876 in honor of Lieut. Mikhail Alexandrovich Pylzov (ca. 1848–1898), member of Nikolai Przhevalsky (also spelled Przewalski and Prjevalsky) expedition to Mongolia, during which holotype was collected

Schizopygopsis scleracanthus (Tsao, Wu, Chen & Zhu 1992) sclero-, from sklērós (Gr. σκληρός), tough or hard; acanthus (L.), from ákantha (Gr. ἄκανθα), thorn, presumably referring to “saw-like thorn” (translation, i.e., osseus ray) on dorsal fin

Schizopygopsis stolickai Steindachner 1866 in honor of Moravian paleontologist Ferdinand Stoliczka (1838–1874), who collected holotype [spelled stoličkai by Steindachner; caron is deleted per ICZN Art. 32.5.2.1 but some authors incorrectly emend spelling to “stoliczkai”]

Schizopygopsis thermalis Herzenstein 1891 Neo-Latin for of or relating to hot springs or water, referring to its habitat at Tan la mountains of Tibet, elevation 4572–4877 m

Schizopygopsis waddellii (Regan 1905) in honor of Lieut.-Col. Laurence Austine Waddell, C.B. [Companion of the Order of the Bath] (1854–1938), British army surgeon, explorer, philologist, linguist, and chemistry and pathology professor, who preserved type specimens in salt before presenting them to the British Museum (Natural History)

Schizopygopsis younghusbandi Regan 1905 patronym not identified but clearly in honor of Lieut.-Col. Francis Edward Younghusband (1863–1942), who led a 1904 British expedition to (and de facto invasion of) Tibet, during which holotype was collected

Schizopygopsis younghusbandi shannaensis Wu, Tsao, Chen & Zhu 1992ensis, Latin suffix denoting place: Shannan Region of Tibet, where it appears to be endemic

Schizopygopsis younghusbandi wui Tchang, Yueh & Hwang 1964 in honor of Wu Hsien-Wen (1900–1985), National Research Institute of Biology, Academia Sinica, for his contributions to Chinese ichthyology