Family SALMONIDAE Jarocki or Schinz 1822 (Salmonids)

Revised 11 Dec. 2024
PDF version (with illustrations and additional information)

Whitefishes
Subfamily COREGONINAE Bonaparte 1845

Subgenus Coregonus

Coregonus Linnaeus 1758 Artedi’s 1738 name for C. lavaretus: core, from kórē (Gr. κόρη), pupil of eye; gōnía (Gr. γωνία), angle or corner, referring to how front part of pupil projects at an acute angle

Coregonus acrinasus Selz, Dönz, Vonlanthen & Seehausen 2020 acer (L.), sharp or pungent; nasus (L.), nose, referring to its pointed snout when viewed in the sagittal plane

Coregonus albellus Fatio 1890 Latinization of the local names Albele and Albuli in Switzerland, derived from albus (L.), white, root of the name Weissfische (whitefish)

Coregonus albula (Linnaeus 1758) Latin for whitish, referring to its silvery coloration

Coregonus alpinus Fatio 1885 Latin for alpine or of the Alps, described from an alpine lake in Switzerland, Lake Thun

Coregonus anaulorum Chereshnev 1996 name coined by Kaganowsky in 1933 but not made available until 1996, etymology not explained, possibly –orum (L.), commemorative suffix, plural, referring to the Anaouls (or Anauls), a sedentary subtribe of the Yukaghir tribe of northern Siberia, who reportedly lived as fishers and hunters in the Anadyr River basin (where this whitefish occurs) in the 17th century

Coregonus arenicolus Kottelat 1997 arena (L.), sand; –colus (L.), dwelling in, referring to its local name, sandfelchen (sand whitefish), along Lake Constance (Konstanz) in Switzerland, Germany and Austria, where it naturally occurs (introduced elsewhere)

Coregonus atterensis Kottelat 1997 ensis, Latin suffix denoting place: Lake Attersee, Austria, where it appears to be endemic

Coregonus austriacus Vogt 1909icus (L.), belonging to: Austria, where it occurs in Lake Attersee (may also occur in Lake Wolfgangsee)

Coregonus baerii Kessler 1864 in honor of Karl Ernst von Baer (1792– 1876), Baltic-German scientist and explorer, who recommended that Kessler join a commission to study the fishes of St. Petersburg Province, Russia, during which this species was discovered

Coregonus baicalensis Dybowski 1874 ensis, Latin suffix denoting place: Russia’s Lake Baikal, where it is endemic

Coregonus baunti Mukhomediyarov 1948 of Baunt Administrative district, home of the Tsipo-Tsipikan lakes, Vitim River system, Siberia, Russia, where this whitefish occurs

Coregonus bavaricus Hofer 1909 icus (L.), belonging to: Upper Bavaria, Germany, where it is endemic to Lake Ammersee

Coregonus bezola Fatio 1888 Latinization of its local name, besoule or bezoule, at Lake Borget, France [extinct; reported by anglers to have disappeared in the 1960s]

Coregonus brienzii Selz, Dönz, Vonlanthen & Seehausen 2020 of Lake Brienz, Switzerland, the only endemic whitefish species known from this lake

Coregonus candidus Goll 1883 Latin for shining white, clear or bright, referring to its pale livery, “almost devoid of pigmentation” (translation), except during the summer, when it may take on a light bluish tint

Coregonus chadary Dybowski 1869 local name for this whitefish (also spelled khadary) in the Burjatskaja Republic of Russia (now the Republic of Buryatia, a federal subject of Russia)

Coregonus clupeaformis (Mitchill 1818) clupea (L.), herring or herring-like fish (but often applied to shad); formis, Neo-Latin scientific adjective of forma (L.), shape or form, described as “more like a shad than any other fish” in general form and color

Coregonus clupeoides Lacepède 1803 -oides, Neo-Latin from eī́dos (Gr. εἶδος), form or shape: clupea, (L.), herring or herring-like fish, referring to its herring-like appearance as demonstrated by its common name in Scotland (type locality) at time of description, “freshwater herring”

Coregonus confusus Fatio 1885 Latin for confused or mixed together, allusion not explained; proposed as a subspecies of C. annectus (=suidteri) from Lake Morat, Switzerland, Fatio tentatively considered rare individuals from nearby lakes Neuchâtel and Bienne as hybrids between it and the local species, which could have hybridized (i.e., mixed together) when the original hydrography and environment of the lakes were modified in 1868–1878

Coregonus danneri Vogt 1908 in honor of fisheries inspector H. Danner (Linz, Austria), who collected holotype

Coregonus duplex Fatio 1890 Latin for double or twofold, allusion not explained, but probably used here to mean duplicitous: proposed as one of four subspecies of C. schinzii (=fera), referring to the “different [taxonomic] assessments” caused by this species’ “misleading forms” (translation)

Coregonus fatioi Kottelat 1997 in honor of Swiss zoologist Victor Fatio (1838–1906), who wrote a standard reference work on Swiss vertebrates (1890) in which he described this species as C. wartmanni alpinus, a name preoccupied by his own C. alpinus (1885)

Coregonus fera Jurine 1825 local name for several species of Coregonus in Switzerland and France, including this one, as recorded by Rondelet in 1555 [extinct due to overfishing and introgression with introduced Coregonus; last seen in 1920]

Coregonus fluviatilis Isachenko 1925 Latin for riverine or of a river, described as a “purely” (translation) river-adapted relative of C. lavaretus and C. pidschian

Coregonus fontanae Schulz & Freyhof 2003 in honor of German poet Theodor Fontane (1819–1898), who published several books reflecting his love for the state of Brandenburg, and dedicated his last book to the people and landscape around Lake Stechlin, Germany, where this whitefish is endemic

Coregonus gutturosus (Gmelin 1818) Latin for goitered or strumose, derived from its German name, Kropf-Maräne (Struma Whitefish), presumably referring to its small head and high body so that throat appears to be hypertrophied with a vague resemblance to a goiter or struma [extinct due to eutrophication; last verifiably seen no later than 1950]

Coregonus heglingus Schinz 1822 Latinization of Hegling, German word for a small kind of whitefish (this species reaches 24 cm SL)

Coregonus hiemalis Jurine 1825 Latin for of winter, referring to its December spawning run from deeper parts of Lake Geneva, Switzerland and France, to shallow areas near shore [extinct due to overfishing and eutrophication; last seen in the early 1900s]

Coregonus hoferi Berg 1932 in honor of German fisheries scientist Bruno Hofer (1861–1916), who reported this species as a variety of C. wartmanni in 1909 [present in the 1940s but probably extinct due to introduction of stocked Coregonus of unknown origin]

Coregonus holsatus Thienemann 1916 Latinization of Holstein (derived from the Northern Low Saxon Holsaten), proposed as a subspecies of C. maraena from Lake Selenter near Lütjenburg, Holstein, Germany

Coregonus intermundia Selz & Seehausen 2023 Latin for “spaces between the worlds,” referring to how this species is intermediate to C. litoralis and C. suspensus in its ecology and in some morphological characters

Coregonus karasjovi Bogdanov 2022 in honor of G. L. Karasjov (romanized as Karasev), who described this species as C. baunti in 1987 but used a preoccupied name (C. baunti Mukhomediyarov 1948)

Coregonus kiletz Michailovsky 1903 local name for this species along Onega Lake of northwestern Russia, derived from the Russian kil, keel, presumably referring to its general body shape

Coregonus ladogae Pravdin, Golubev & Belyaeva 1938 of Lake Ladoga, Russia, where it is endemic (but introduced into other lakes)

Coregonus lavaretus (Linnaeus 1758) Latinization of lavaret, its vernacular name in the Savoyard dialect of France and Switzerland, as recorded by Rondelet in 1555

Coregonus litoralis Selz & Seehausen 2023 Latin for “coming from the shore,” referring to its spawning habitat, in very shallow water of a few meters close to or directly at the lake shore

Coregonus lucinensis Thienemann 1933ensis, Latin suffix denoting place: Lake Breiter Lucin (or Luzin), Mecklenburg-Strelitz, Germany, where it is endemic

Coregonus lutokka Kottelat, Bogutskaya & Freyhof 2005 Finnish variant of ludoga, known as the Ludoga Whitefish, referring to its occurrence in Lake Ladoga (sometimes spelled Ludoga), Russia [replacement name for Coregonus widegreeni ludoga Berg 1916, preoccupied by Coregonus ludoga Polyakov 1874 (=C. widegreni)]

Coregonus macrophthalmus Nüsslin 1882 big-eyed, from makrós (Gr. μακρός), long or large, and ophthalmós (Gr. ὀφθαλμός), eye, referring to its very large eyes, their diameter representing at least 5% of body length

Coregonus maraena (Bloch 1779) Latinization of Maräne, German word for whitefish

Coregonus maraenoides Polyakov 1874 -oides, Neo-Latin from eī́dos (Gr. εἶδος), form or shape: referring to its similarity to and previous identification as C. maraena

Coregonus maxillaris Günther 1866 Latin for of or belonging to the jaw, referring to its “well developed, strong” maxillary

Coregonus megalops Widegren 1863 mégas (Gr. μέγας), big; ṓps (Gr. ὦψ), eye, referring to large eye, its diameter just over 1/3 length of head

Coregonus muelleri Selz & Seehausen 2023 in honor of the late Rudolf Müller (1944–2023), fisheries biologist, Center for Ecology, Evolution and Biogeochemistry (Kastanienbaum, Switzerland), who dedicated a large part of his career to understanding the ecology of whitefishes in Swiss lakes, and confirmed the rediscovery of C. nobilis

Coregonus muksun (Pallas 1814) Muksûn, its local Russian name in Siberia

Coregonus nasus (Pallas 1776) Latin for nose, referring to its rounded snout

Coregonus nelsonii Bean 1884 in honor of American naturalist and ethnologist Edward W. Nelson (1855–1934), who collected holotype, for his “important zoological researches” in Alaska

Coregonus nilssoni Valenciennes 1848 in honor of Swedish zoologist and archaeologist Sven Nilsson (1787–1883), whose 1832 work on Scandinavian fishes is cited several times by Valenciennes

Coregonus nobilis Haack 1882 Latin for well-known, noted or celebrated, allusion not explained, perhaps referring to how it was “often caught in enormous quantities” (translation) during the spawning season in Lake Vierwaldstätter, Switzerland, where it is endemic

Coregonus obliterus Selz & Seehausen 2023 Latin for forgotten, this whitefish having been forgotten for more than a century in the scientific literature [extinct by 1935 due to eutrophication]

Coregonus oxyrinchus (Linnaeus 1758) sharp-snouted, from oxýs (Gr. ὀξύς), sharp or pointed, and rhýnchos (Gr. ῥύγχος), snout, referring to its long, pointed snout [possibly a synonym of C. lavaretus]

Coregonus palaea Cuvier 1829 presumably a Latinization of palée, its local name in Switzerland, where it is endemic

Coregonus pallasii Valenciennes 1848 in honor of Prussian naturalist and explorer Peter Simon Pallas (1741–1811), whose collection provided holotype, with “respect for the memory of this great naturalist” (translation)

Coregonus pennantii Valenciennes 1848 in honor of Welsh naturalist and writer Thomas Pennant (1726–1798), who wrote about this species (called the “Gwiniad”) in 1769 but referred to it as C. lavaretus

Coregonus pidschian (Gmelin 1789) local name for this species in the Ob River region of Siberia, its presumed type locality

Coregonus pollan Thompson 1835 name by which it is “invariably known” at Lough Neah, Ireland, type locality

Coregonus pravdinellus Dulkeit 1950 ellus (L.), diminutive connoting endearment: in honor of Soviet ichthyologist Ivan Fedorovich Pravdin (1880–1963), who co-described Coregonus ladogae in 1938

Coregonus profundus Selz, Dönz, Vonlanthen & Seehausen 2020 Latin for deep, referring to its unique ecology of living and breeding in great depths in Lake Thun (Switzerland)

Coregonus renke (Schrank 1783) local name for this species at Lake Starnberg, Germany, type locality

Coregonus restrictus Fatio 1885 Latin for confined, allusion not explained, perhaps referring to its being confined to Lake Morat, Switzerland, where it is now extinct

Coregonus sajanensis Gundriser 1978ensis, Latin suffix denoting place: Sajany (romanized as Sayan) Mountains, which overlook the Todzha depression in Tyva, Russia, where this species was reported in 1966

Coregonus sarnensis Selz & Seehausen 2023ensis, Latin suffix denoting place: Sarnen, Switzerland, a village on the shores of Lake Sarnen, type locality

Coregonus skrjabini Karasev 1987 in honor of Anatoly Georgievich Scryabin (1938–2000), for his contribution to the study of whitefishes from the Baunt Lakes system

Coregonus smitti Warpachowski 1900 patronym not identified but clearly in honor of Swedish zoologist Fredrik Adam Smitt (1839–1904), whose 1883 revision of Coregonus and system of measuring coregonine fishes are cited by Warpachowski

Coregonus steinmanni Selz, Dönz, Vonlanthen & Seehausen 2020 in honor of Swiss biologist Paul Steinmann (1885–1953), whose 1950 monograph on Swiss Coregonus is the “most comprehensive compendium” on the subject to date, and whose large collection of Swiss fishes has been “essential to describe the whitefish diversity that was present in Switzerland just before or at the beginning of the strong anthropogenic-induced eutrophication of many Swiss lakes which was accompanied by population collapse, speciation reversals, and extinction of Swiss whitefish”

Coregonus stigmaticus Regan 1908 Latin for marked, referring to small blackish spots on sides

Coregonus suidteri Fatio 1885 patronym not explained but almost certainly in honor of Otto Suidter (1833–1901), a Swiss pharmacist and naturalist who studied coregonine fishes [extinct due to eutrophication; last seen 1899]

Coregonus supersum Selz & Seehausen 2023 Latin for to be over and above, be left or remain (or “I survive” or “I am left over” per the authors) the only surviving whitefish species in Lake Zug, Switzerland

Coregonus suspensus Selz & Seehausen 2023 Latin for suspended or levitating, referring to its only being caught during the spawning season in the pelagic water column

Coregonus trybomi Svärdson 1979 in honor of “prominent” Swedish fisheries biologist Filip Trybom (1850–1913), who was the first to publish on the existence of this species and study its morphology in 1903

Coregonus vandesius Richardson 1836 Latinization of vendace, local name for this species at Lochmaben, Scotland, type locality

Coregonus vessicus Dryagin 1932 icus (L.), belonging to: Ves’, old Russian name for Vespians, Finnish people who lived on the shores of Lake Belozero, Karelia, Russia, type locality

Coregonus wartmanni (Bloch 1784) in honor of Swiss physician-naturalist Bernhard Wartmann (1739-1815), who described this species in 1777 but did not use a Linnaean name

Coregonus widegreni Malmgren 1863 in honor of Swedish ichthyologist and fisheries official Hjalmar Widegren (1838-1878), who recognized this species as a unique form of C. fera in 1862

Coregonus zuerichensis Nüsslin 1882 ensis, Latin suffix denoting place: Lake Zürich, Switzerland, type locality

Coregonus zugensis Nüsslin 1882 ensis, Latin suffix denoting place: Lake Zug, Switzerland, type locality [extinct by 1935 due to eutrophication]

Coregonus (subgenus Leucichthys) Dybowski 1874 leukós (Gr. λευκός), white; ichthýs (Gr. ἰχθύς), fish, Greek translation of the Anglo-Saxon vernacular “whitefish,” referring to their generally white or silvery coloration

Coregonus alpenae (Koelz 1924) of Alpena, Michigan, USA, a city on the shore of Lake Michigan’s Thunder Bay, one of the locations where this whitefish was collected (but not considered part of the paratype series)

Coregonus artedi Lesueur 1818 patronym not identified but almost certainly in honor of Swedish naturalist Peter Artedi (1705–1735), often described as the “father of ichthyology” [presumably a noun in apposition, without the genitive “i”]

Coregonus artedi hubbsi (Koelz 1929) in honor of American ichthyologist Carl L. Hubbs (1894–1979), who conducted “extensive explorations” of the eight lakes of the Huron Mountain Club in Marquette County, Michigan, USA, and whose specimens were used in this description

Coregonus autumnalis (Pallas 1776) Latin for autumnal, referring to how it migrates from the sea to fresh water in “immense numbers” (translation) during the autumn

Coregonus hoyi (Milner 1874) in honor of American physician-naturalist Philo R. Hoy (1816–1892), Racine, Wisconsin, USA, who provided holotype

Coregonus johannae (Wagner 1910) in honor of Johanna, “a slight token of gratitude for my great indebtedness to my life-companion” (no other information available)

Coregonus kiyi (Koelz 1921) name for this species used by the commercial fishermen of Lake Michigan (rhymes with eye-eye)

Coregonus kiyi orientalis (Koelz 1929) Latin for eastern, referring to Lake Ontario, the easternmost of the Great Lakes (where it was endemic and last seen in 1964, extinct due to commercial overexploitation and introduced species)

Coregonus laurettae Bean 1881 in honor of Bean’s wife, Lauretta

Coregonus migratorius (Georgi 1775) Latin for migratory, referring to its spawning migrations into rivers, which Georgi noted did not seem to follow a set pattern year after year (e.g., different times of years and types of rivers)

Coregonus nigripinnis (Milner 1874) niger (L.), dark or black; pinnis, Neo-Latin adjective of pinna (L.), fin, i.e., finned, referring to black outer halves of fins [extinct due to commercial overexploitation and introduced species; last seen in 1969 but still may be extant in Lake Nipigon as C. n. regalis]

Coregonus nigripinnis regalis (Koelz 1929) Latin for royal, allusion not explained nor evident

Coregonus nipigon (Koelz 1925) named for Lake Nipigon (Ontario, Canada), type locality; presumably derived from the Ojibwe word Animbiigoong, meaning “at continuous water” or “at waters that extend [over the horizon]”

Coregonus peled (Gmelin 1789) presumably from pelyad, its local name along the Pechora River in northern Russia, type locality

Coregonus reighardi (Koelz 1924) in honor of American zoologist Jacob Reighard (1861–1942), Koelz’ professor at the University of Michigan and director of the biological survey of the Great Lakes under the United States Fish Commission [probably extinct, last seen in 1985]

Coregonus sardinella Valenciennes 1848 diminutive of Sardina, a sardine, referring to its herring-like shape [treated as a synonym of C. albula by some workers]

Coregonus subautumnalis Kaganowsky 1932 etymology not explained, presumably sub, (L.), less or under (i.e., somewhat), referring to its close relationship with C. autumnalis

Coregonus tugun (Pallas 1814) Tughùn, its local Russian name in Siberia

Coregonus ussuriensis Berg 1906ensis, Latin suffix denoting place: Ussuri River, Russia, where part of the type series was collected

Coregonus zenithicus (Jordan & Evermann 1909)icus (L.), belonging to: Duluth, Minnesota, USA, the “Zenith City,” where Jordan and Evermann saw hundreds of this species in a cold-storage plant

Coregonus zenithicus bartletti (Koelz 1931) in honor of Harley Harris Bartlett (1886–1960), American botanist, ethnographer, collector, and Koelz’ colleague at the University of Michigan

Coregonus (subgenus Incertae sedis)

Coregonus huntsmani Scott 1987 in honor of Canadian marine biologist Archibald Gowalock Huntsman (1883–1973), who recognized the species as distinct as early as 1921 [replacement name for C. canadensis Scott 1967, preoccupied by C. nasus canadensis Berg 1932]

Prosopium Jordan 1878 from prosōpeī́on (Gr. προσωπεῖον), mask, referring to the large bones (preorbitals) in front of eyes of P. quadrilateralis (=cylindraceum)

Prosopium abyssicola (Snyder 1919) abyssus (L.), from ábyssos (Gr. ἄβυσσος), the deep sea; –cola (L.), dweller or inhabitant, referring to its deep-water habitat in Bear Lake, Utah-Idaho border, USA (Snyder also noted that it spawned late winter at a depth of 30 m)

Prosopium coulterii (Eigenmann & Eigenmann 1892) in honor of American botanist John Merle Coulter (1851–1928), author of Manual of the Botany of the Rocky Mountain Region (the area where this species occurs)

Prosopium cylindraceum (Pennant 1784) Latin for cylindrical, referring to its “almost cylindrical” body

Prosopium gemmifer (Snyder 1919) Latin for producing gems, probably referring to its “pearly” breeding tubercles

Prosopium spilonotus (Snyder 1919) spílos (Gr. σπίλος), mark or spot; nótos (Gr. νότος), back, referring to round, dusky spots on dorsal region (which disappear with age)

Prosopium williamsoni (Girard 1856) in honor of Lt. Robert S. Williamson (1825–1882), U.S. Army Corps of Topographical Engineers, who commanded a government survey of proposed transcontinental railroad routes, during which holotype was collected

Stenodus Richardson 1836 sténos (Gr. στένος), narrow; odoús (Gr. ὀδούς), tooth, referring to narrow bands of minute teeth on both jaws

Stenodus leucichthys (Güldenstädt 1772) leukós (Gr. λευκός), white; ichthýs (Gr. ἰχθύς), fish, Greek translation of Belaia Rybyza, its Russian vernacular name

Stenodus nelma (Pallas 1773) local Russian name for this species in Siberia


Graylings
Subfamily THYMALLINAE Gill 1885

Thymallus Linck 1790 presumably tautonymous with Salmo thymallus Linnaeus 1758 (see T. thymallus for etymology)

Thymallus aeliani (Valenciennes 1848) in honor of Roman author and naturalist Claudius Aelianus (ca. 175–ca. 235), who wrote about this species (or one similar to it) in his De Natura Animalium

Thymallus arcticus (Pallas 1776)icus (L.), belonging to: Arctic Ocean basin of Siberia, Russia, where it occurs in “rocky streams”

Thymallus baicalensis Dybowski 1874 ensis, Latin suffix denoting place: described as a Lake Baikal (Russia) variety of T. grubi (also occurs in Mongolia)

Thymallus baicalolenensis Matveyev, Samusenok, Pronin & Tel’pukhovsky 2005ensis, Latin suffix denoting place: baicol and lena, referring to distribution in Lake Baikal basin and Lena River, Siberia, respectively

Thymallus brevicephalis Mitrofanov 1971 short-headed, from brevis (L.), short, and kephalḗ (Gr. κεφαλή), head, referring to shorter and lower head relative to body length compared with T. arcticus [originally spelled brevicephalis, often incorrectly emended to brevicephalus]

Thymallus brevipinnis Svetovidov 1931 brevis (L.), short; pinnis, Neo-Latin adjective of pinna (L.), fin, referring to shorter dorsal and anal fins compared with T. arcticus

Thymallus brevirostris Kessler 1879 brevis (L.), short; rostris, Neo-Latin scientific adjective of rostrum (L.), snout, referring to its “noticeably shorter” (translation) snout compared with T. vulgaris (=thymallus)

Thymallus burejensis Antonov 2004ensis, Latin suffix denoting place: Bureye (or Bureya) River, tributary of Middle Amur River, Russia, where it is endemic

Thymallus flavomaculatus Knizhin, Antonov & Weiss 2006 flavus (L.), yellow; maculatus (L.), spotted, referring to orange-yellow spots on dorsal fin

Thymallus grubii Dybowski 1869 patronym not identified, probably in honor of Dybowski’s professor, the Prussian zoologist Adolph Eduard Grube (1812–1880)

Thymallus lenensis Knizhin 2022ensis, Latin suffix denoting place: upper Lena River basin, Russia, where it primarily occurs

Thymallus ligericus Persat, Weiss, Froufe, Secci-Petretto & Denys 2019icus (L.), belonging to: Liger, Latin name for Loire River basin (France), where it is endemic

Thymallus mertensii Valenciennes 1848 in honor of German naturalist and explorer Karl Heinrich Mertens (1796–1830), who illustrated the plate from which Valenciennes proposed the species

Thymallus nigrescens Dorogostaisky 1923 Latin for blackish, referring to dark (not mottled) coloration, blackish-blue or almost black during spawning

Thymallus nikolskyi Kaschenko 1899 in honor of Russian ichthyologist-herpetologist Alexander Mikhailovich Nikolsky (also spelled Nikolskii, 1858–1942), who, with Solomon Markovich Herzenstein (1854–1894), first indicated differences between Altai and European graylings

Thymallus pallasii Valenciennes 1848 in honor of Prussian naturalist and explorer Peter Simon Pallas (1741–1811), who described T. arcticus in 1776

Thymallus signifer (Richardson 1823) signum (L.), flag or banner; –fer, from fero (L.), to have or bear, referring to its large dorsal fin, whose rays increase “rapidly in height, as their origin is more posterior, become more and more branched, and cause the fin to play loosely like a flag over the posterior part of the body”

Thymallus svetovidovi Knizhin & Weiss 2009 in honor of Russian ichthyologist Anatolii Nikolaevich Svetovidov (1903–1985), “famous researcher of graylings of Eurasia” (translation)

Thymallus thymallus (Linnaeus 1758) thýmallos (Gr. θύμαλλος), named for how it smells of freshly plucked Thymus, or thyme, as reported by Roman author Claudius Aelianus (see T. aeliani, above); whether the fish actually smells of thyme is a matter of some debate among anglers, naturalist and ichthyologists

Thymallus tugarinae Knizhin, Antonov, Safronov & Weiss 2007 in honor of Polina Yakovlevna Tugarina (1928–2004), Irkutsk State University, “renowned” (translation) researcher of the graylings of Siberia and the Russian Far East

Thymallus yaluensis Mori 1928ensis, Latin suffix denoting place: upper Yalu River at Kozan, Korea, type locality [sometimes spelled jaluensis]


Trouts, Salmons and Chars
Subfamily SALMONINAE Jarocki or Schinz 1822

Brachymystax Günther 1866 brachýs (Gr. βραχύς), short; mýstax (Gr. μύσταξ), upper lip or moustache, referring to “rather short” maxillary of B. coregonoides (=lenok)

Brachymystax czerskii Kirillov 1979 patronym not identified, probably in honor of Jan Czerski (1845–1892), Polish paleontologist, geologist, geographer and explorer of Siberia, where this species occurs

Brachymystax lenok (Pallas 1773) Lénok, its local Russian name in Siberia

Brachymystax savinovi Mitrofanov 1959 patronym not identified, possibly mammalogist E. F. Savinov, one of Mitrofanov’s colleagues at the Academy of Sciences Kazakhstan USSR, Institute of Zoology

Brachymystax tsinlingensis Li 1966ensis, Latin suffix denoting place: Tsinling Mountain region, Shaanxi Province, China, type locality

Brachymystax tumensis Mori 1930ensis, Latin suffix denoting place: Tumen (also spelled Tuman) River, Yen-gan, Korea, type locality

Hucho Günther 1866 tautonymous with Salmo hucho Linnaeus 1758, presumably a Latinization of huchen, German name for H. hucho

Hucho bleekeri Kimura 1934 in honor of Dutch army surgeon and ichthyologist Pieter Bleeker (1819–1878), who described the “allied” Salmo pomatops (species inquirenda) in 1879

Hucho hucho (Linnaeus 1758) presumably a Latinization of huchen, German name for this species

Hucho ishikawae Mori 1928 in honor of Japanese biologist Chiyomatsu Ishikawa (1861–1935), Tokyo Imperial University

Hucho taimen (Pallas 1773) Taïmen, its local Russian name in Siberia

Oncorhynchus Suckley 1861 ónkos (Gr. ὄγκος), hook; rhynchus, from rhýnchos (Gr. ῥύγχος), snout (i.e., jaw), referring to hooked lower jaw, or kype, of breeding males

Oncorhynchus gorbuscha (Walbaum 1792) derived from gorbúša, Russian word for humpback, referring to the pronounced humped back of males during their spawning migration

Oncorhynchus gorbuschka Glubokovsky & Zhivotovsky 2024 same name as O. gorbuscha but differing by one letter, emphasizing their “crypticity and close phylogenetic relationship” (this species spawns in odd- rather than even-numbered years)

Oncorhynchus kawamurae Jordan & McGregor 1925 in honor of Japanese biologist-limnologist Tamiji Kawamura (1883–1964), Imperial University (Kyoto, Japan), who “presented” holotype [although named after a man, “ae” is an acceptable way to form a genitive from a masculine noun that ends in “a”]

Oncorhynchus keta (Walbaum 1792) local name for this species in the Evenki language of eastern Siberia

Oncorhynchus kisutch (Walbaum 1792) from kysutch (pronounced keez-utch), vernacular name for this species in the Kamchatka Peninsula in the 16th century (Walbaum spelled it variously as hisutch, hisatch and kisatch; although kisutch is the younger name, prevailing usage applies since 1899)

Oncorhynchus masou (Brevoort 1856) from masu, vernacular name for this species in Japan

Oncorhynchus masou formosanus (Jordan & Oshima 1919)anus (L.), belonging to: Formosa (Taiwan), where it is endemic

Oncorhynchus masou ishikawae Jordan & McGregor 1925 in honor of Japanese biologist Chiyomatsu Ishikawa (1861–1935), Imperial Museum of Tokyo, who collected some of the salmonid specimens examined by the authors

Oncorhynchus masou macrostomus (Günther 1877) big-mouthed, from makrós (Gr. μακρός), long or large, and stóma (Gr. στόμα), mouth, referring to its “wide oblique” mouth

Oncorhynchus masou rhodurus Jordan & McGregor 1925 rosy-tailed, from rhódon (Gr. ῥόδον), and ourá (Gr. οὐρά), tail, presumably referring to a bright red stripe along outer rays of caudal fin (fading to white in spirits)

Oncorhynchus nerka (Walbaum 1792) vernacular name (also spelled narka) for this salmon used in the Kamchatka Peninsula in the 16th century

Oncorhynchus tshawytscha (Walbaum 1792) derived from chavycha (pronounced cha-vee-cha or cho-wee-cha), vernacular name for this salmon used in the Kamchatka Peninsula in the 16th century

Oncorhynchus (subgenus Parasalmo) Vladykov 1972 pará (Gr. παρά), near, for Pacific-area salmonids then placed in Salmo [proposed by Vladykov in 1963 but not available until 1972]

Oncorhynchus aguabonita (Jordan 1892) agua and bonita, Spanish for water and beautiful, respectively, named for Agua Bonita Falls, the cataract (or cascade) at Volcano Creek, California, USA, near type locality

Oncorhynchus aguabonita gilberti (Jordan 1894) in honor of American ichthyologist and fisheries biologist Charles H. Gilbert (1859-1928) who collected holotype

Oncorhynchus aguabonita whitei (Evermann 1906) in honor of American adventurer and nature writer Stewart Edward White (1873–1946), who, fearing the overfishing and potential extinction of California’s golden trouts (of which this is one), contacted his friend and fellow outdoors enthusiast U.S. president Theodore Roosevelt, who then ordered the government to research their biology, distribution and propagation (with Evermann as the lead investigator)

Oncorhynchus apache (Miller 1972) “… in view of the deep concern of the [White Mountain] Apache Tribe for the welfare of this species, it is most appropriate that this distinctive fish be named”

Oncorhynchus chrysogaster (Needham & Gard 1964) chrysós (Gr. χρυσός), gold or golden; gastḗr (Gr. γαστήρ), belly or stomach, referring to bright-orange color on belly

Oncorhynchus clarkii (Richardson 1836) in “tribute to the memory” of explorer William Clark (1770–1838, misspelled Clarke by Richardson), who, with Meriwether Lewis (see O. lewisi), caught this trout during the Lewis and Clark (or Corps of Discovery) Expedition (1804–06), the first American expedition to cross what is now the western portion of the United States

Oncorhynchus gilae (Miller 1950) of the Upper Gila River system, New Mexico, where this trout has “managed to persist” (Arizona populations are now extirpated)

Oncorhynchus henshawi (Gill & Jordan 1878) in honor of American ornithologist Henry W. Henshaw (1850–1930), U.S. Geological Survey, who collected holotype

Oncorhynchus henshawi alvordensis (Sigler & Sigler 1987)ensis, Latin suffix denoting place: Alvord Basin, Oregon and Nevada, USA, where it occurred [extinct due to hybridization with stocked O. mykiss in the 1920s]

Oncorhynchus henshawi humboldtensis Trotter & Behnke 2008 ensis, Latin suffix denoting place: Humboldt River drainage, Nevada and Oregon, USA, where it is endemic

Oncorhynchus henshawi seleniris (Snyder 1933) combination of Selene and Iris, Greek goddesses of the moon and rainbow, respectively, allusion not explained but according to Snyder’s 1908 description of the ribbonfish Trachypterus seleniris (=Trachipterus altivelis), which he named “on account of a fancied resemblance of the long, flat, silvery body to the colorless lunar rainbow,” referring to this trout’s pale coloration, “the whole body much suffused with yellow,” and absence of spots

Oncorhynchus lewisi (Girard 1856) in honor of American explorer Meriwether Lewis (1774–1809), who, with William Clark (see above), caught six of this trout at Great Falls, on the Missouri River, Montana, during the Lewis and Clark (or Corps of Discovery) Expedition (1804–06), the first American expedition to cross what is now the western portion of the United States

Oncorhynchus mykiss (Walbaum 1792) from mykizha, vernacular name for this species used in the Kamchatka Peninsula in the 16th century

Oncorhynchus mykiss aquilarum (Snyder 1917) genitive plural of aquila (L.), eagle, referring to Eagle Lake, California, USA, where it is endemic

Oncorhynchus mykiss calisulat Campbell & Conway 2023 from the Winnemem Wintu name for this subspecies, cali sulat, meaning “beautiful trout” (the Winnemem Wintu are a Native American tribe who were originally located along the McCloud River, Shasta County, California, where this trout occurs)

Oncorhynchus mykiss gairdnerii (Richardson 1836) in honor of British surgeon-naturalist Meredith Gairdner (1809–1837), Hudson’s Bay Company (a fur trading company), who collected type specimens and sent them to Richardson, along with a detailed written description which Richardson quoted; name is “intended as a tribute to the merits of a young though able naturalist, from whom science may expect many important acquisitions, and especially in the history of the Zoology of the Northwest coast of America, should his engagements with the Hudson’s Bay Company permit him to cultivate that hitherto neglected field of observation” (Gairdner died a year later from tuberculosis)

Oncorhynchus mykiss irideus (Gibbons 1855) Latin for rainbow-like or iridescent, referring to its coloration, a fitting description for a fish that would later be known as the Rainbow Trout

Oncorhynchus mykiss nelsoni (Evermann 1908) in honor of American naturalist and ethnologist Edward W. Nelson (1855–1934), who collected holotype

Oncorhynchus mykiss newberrii (Girard 1859) in honor of John Strong Newberry (1822–1892), American geologist, physician and explorer, who collected holotype

Oncorhynchus mykiss stonei (Jordan 1894) in honor of Livingston Stone (1836–1913), superintendent of the first U.S. fish hatchery (The Baird Hatchery, Shasta County, California), who collected holotype at or near the hatchery; Jordan & Evermann (1896) said name honors Stone for his “valuable services in the propagation of salmon and trout”

Oncorhynchus penshinensis (Pallas 1814)ensis, Latin suffix denoting place: Gulf of Penshine, Norofskuja River, Russia, type locality

Oncorhynchus virginalis (Girard 1856) Latin for maidenly, allusion not explained; according to James Prosek’s Trout: an Illustrated History (1996), “its qualities suited a maiden: a pure, unsullied beauty, delicate and desirable”

Oncorhynchus virginalis behnkei Montgomery 1995 in honor of salmonid taxonomist Robert J. Behnke (1929–2013) [Montgomery, a Boston Globe columnist, reprinted descriptive information from Behnke (1992) under the name O. clarkii behnkei in a book about fly fishing (Many Rivers to Cross), making him the author of the taxon]

Oncorhynchus virginalis bouvieri (Jordan & Gilbert 1883) name proposed by U.S. Army officer and ornithologist Charles E. Bendire (1836–1897), in honor of his friend “Captain Bouvier,” possibly John Vernou Bouvier, Sr. (1843–1926), 80th New York Infantry during the U.S. Civil War

Oncorhynchus virginalis macdonaldi (Jordan & Evermann 1890) in honor of Marhsall McDonald (1835–1895, note Latinization of “Mc” to “Mac”), U.S. Commissioner of Fisheries, for his services in spreading the range of salmonids in America [extinct due to introgressive hybridization with stocked O. mykiss; last seen ca. 1910]

Oncorhynchus virginalis pleuriticus (Cope 1872)icus (L.), pertaining to: pleuri-, from pleurá (Gr. πλευρά), side, referring to its red lateral band

Oncorhynchus virginalis stomias (Cope 1871) stomías (Gr. στομίας), a hard-mouthed horse but often used for any large-mouthed animal, referring to its “wide mandible and mouth” (although its mouth is no larger than other cuthroat trout subspecies) [this taxon is subject to revalidation since holotype is actually a specimen of O. v. virginalis]

Oncorhynchus virginalis utah (Suckley 1874) named for Utah Lake, Utah, USA, type locality

Parahucho Vladykov 1963 pará (Gr. παρά), proposed as a subgenus of Hucho

Parahucho perryi (Brevoort 1856) in honor of Matthew C. Perry (1794 –1858), Commodore, U.S. Navy, “able commander of the United States Japan Expedition, to whose efforts alone we owe the scanty yet interesting zoological collections and drawings [including that of this species], made under disadvantageous circumstances, while the squadron was in those distant seas”

Salmo Linnaeus 1758 Latin for trout or salmon

Salmo abanticus Tortonese 1954icus (L.), belonging to: Lake Abant, Turkey, type locality

Salmo akairos Delling & Doadrio 2005 ákairos (Gr. άκαιρος), inopportune, allusion not explained in description; per Delling (pers. comm.), name reflects the set of circumstances that prevented his first attempt to describe this species (with a different name) from being published in 2002 (rejection by journal, unauthorized release of manuscript name to the popular press, and a general reticence to describe new Salmo taxa due to prevailing notions of salmonid taxonomy)

Salmo aphelios Kottelat 1997 ápo (Gr. άπω), far away or remote; hḗlios (Gr. ἥλιος), sun, i.e., aphelion, astronomic term for point of orbit where a planet is at its greatest distance from the sun (early July for Earth), referring to its June-July spawning time [replacement name for Salmo letnica aestivalis Stefanovic 1948, preoccupied by Salmo ischchan aestivalis Fortunatov 1927]

Salmo araxensis Turan, Kottelat & Kaya 2022ensis, Latin suffix denoting place: Arax, ancient name of the Aras River, Kars Province, Turkey, in whose basin this trout occurs

Salmo ardahanensis Turan, Kottelat & Kaya 2022ensis, Latin suffix denoting place: Ardahan Province, Turkey, where type locality (Stream Toros, Kura River drainage) is situated

Salmo balcanicus (Karaman 1927)icus (L.), belonging to: the Balkans, referring to its distribution in Lake Ohrid (Albania and Macedonia), the deepest lake of the Balkans

Salmo baliki Turan, Aksu, Oral, Kaya & Bayçelebi 2021 in honor of Turkish ichthyologist Süleyman Balik, for his contributions to the freshwater fish fauna of Turkey

Salmo brunoi Turan, Bayçelebi, Aksu & Oral 2024 in honor of Bruno Guinand, University of Montpellier (France), for his “valuable contribution” to Salmo population genomics research

Salmo carpio Linnaeus 1758 Latin for carp but in this case probably referring to carpione, this trout’s vernacular name in Italy (where it is native to Lake Garda), as recorded in Salviani’s Aquatilium animalium (1558)

Salmo caspius Kessler 1877 Latin for Caspian, referring to is distribution in the Caspian Sea drainage basin

Salmo cenerinus Nardo 1847 apparent Latinization of the Italian cenere, ashen or ashy, name coined by Italian malacologist Stefano Abate Chiereghini (1745–1820) in an unpublished manuscript, where he described this trout as having “shiny silver ashy [sides] sown with circular dark red spots” (translation)

Salmo cettii Rafinesque 1810 in honor of Francesco Cetti (1726–1778), Jesuit priest, zoologist and mathematician, who referred to this species as “Trotta Sonda” in his multi-volume natural history of Sardinia (1774–77)

Salmo chilo Turan, Kottelat & Engin 2012 Latin word for someone who has a big lip, referring to its “conspicuously” subterminal mouth with a fleshy maxilla and lower lip

Salmo ciscaucasicus Dorofeeva 1967 -icus (L.), belonging to: Ciscaucasia, referring to distribution in northeast Caucasus (between Black and Caspian Seas)

Salmo coruhensis Turan, Kottelat & Engin 2010ensis, Latin suffix denoting place: Çoruh River drainage, Erzurum Province, Turkey, type locality

Salmo danilewskii (Gulelmi 1888) patronym not identified, presumably in honor of Nikolai Danilewski (also spelled Nikolay Danilevsky, 1822–1885), Russian naturalist, economist, ethnologist, philosopher and historian

Salmo dentex (Heckel 1851) from dens (L.), tooth, referring to its larger, more powerful teeth compared with typical S. trutta in Dalmatia (now Bosnia-Herzegovina)

Salmo duhani Turan & Aksu 2023 in honor of Duhan Turan, son of the first author

Salmo ekmekciae Küçük, Kalaycı, Güçlü, Oral & Turan 2024 in honor of Turkish ichthyologist Fitnat Güler Ekmekçi, Hacettepe University, “in appreciation of her contribution to literature”

Salmo euphrataeus Turan, Kottelat & Engin 2014eus (L.), adjectival suffix: headwaters of the northern Euphrates River in Turkey, where it occurs

Salmo ezenami Berg 1948 of Lake Ezenam, Daghestan, Caucasus, Russia, type locality

Salmo fahrettini Turan, Kalayci, Bektaş, Kaya & Bayçelebi 2020 in honor of Turkish zoologist Fahrettin Küçük, Süleyman Demirel University, for his contributions to the knowledge of freshwater fishes in Turkey (where this trout is endemic)

Salmo farioides Karaman 1938 -oides, Neo-Latin from eī́dos (Gr. εἶδος), form or shape: referring to its previous identification under the name S. fario (=trutta)

Salmo ferox Jardine 1835 Latin for fierce, referring to its “extreme voracity and rapacious habits”

Salmo fibreni Zerunian & Gandolfi 1990 of the Fibreno river basin, including Lake Posta Fibreno (Italy), where it is endemic

Salmo gegarkuni Kessler 1877 historical name for Lake Sevan, Armenia (also spelled Gegharkuni), where this species is endemic (although introduced elsewhere)

Salmo ghigii Pomini 1941 in honor of Italian zoologist Alessandro Ghigi (1875–1970), who led expedition during which holotype (now lost) was collected

Salmo ischchan Kessler 1877 Armenian for prince, local name for this trout in Armenia, referring to row of spots like a crown on its head

Salmo ischchan aestivalis Fortunatov 1926 Latin for of the summer, referring to the timing of its spawning run

Salmo kottelati Turan, Doğan, Kaya & Kanyılmaz 2014 in honor of Swiss ichthyologist Maurice Kottelat (b. 1957), for his contributions to the knowledge of European and Asian fishes

Salmo labecula Turan, Kottelat & Engin 2012 Latin for a slight stain or blemish (authors say “with small spots”), referring to black-spotted color pattern of adults

Salmo labrax Pallas 1814 lábrax (Gr. λάβραξ), derived from lavraki, name used by the Crimean Greeks for the anadromous trout of the Black Sea (in Greece, the European Bass Dicentrarchus labrax is also called lavraki)

Salmo letnica (Karaman 1924) local Slavic word meaning “summer fish” because it is caught mainly in the summer, compared with S. dentex, “zimnica,” which is caught mainly in the winter

Salmo lourosensis Delling 2011ensis, Latin suffix denoting place: Louros River, Greece, where it is endemic

Salmo lumi Poljakov 1958 Albanian word for river, proposed as a river-spawning subspecies of the lacustrine S. letnica

Salmo macedonicus (Karaman 1924)icus (L.), belonging to: Macedonia, where type locality (mouth of Treska River, Vardar basin) is situated

Salmo macrostigma (Duméril 1858) macro-, from makrós (Gr. μaκρóς), long or large; stígma (Gr. στίγμα), mark or spot, referring to many black rounded spots regularly arranged on the sides

Salmo marmoratus Cuvier 1829 Latin for marbled, referring to its distinctive color pattern

Salmo multipunctatus Doadrio, Perea & Yahyaoui 2015 multi– (L.), many; punctatus (L.), spotted, referring to numerous spots and marks along body

Salmo munzuricus Turan, Kottelat & Kaya 2017 ensis, Latin suffix denoting place: Munzur Stream, Euphrates River drainage, Turkey, type locality

Salmo murathani Turan, Kottelat & Kaya 2022 in honor of Murathan Turan, son of the first author

Salmo nigripinnis Günther 1866 niger (L.), dark or black; pinnis, Neo-Latin adjective of pinna (L.), fin, i.e., finned, referring to black spots on dorsal fin, black vertical fins, and/or paired fins whose outer halves are black

Salmo okumusi Turan, Kottelat & Engin 2014 in honor of the late Ibrahim Okumuş (1960–2009), Dean of the Faculty of Fisheries, Rize University (Turkey)

Salmo opimus Turan, Kottelat & Engin 2012 Latin for fat, plump or opulent, referring to “massive, deep body” of large adults

Salmo oxianus Kessler 1874anus (L.), belonging to: Oxua River, ancient name for Amu Darya River, referring to distribution in the Upper Amu Darya River basin

Salmo pallaryi Pellegrin 1924 in honor of French-Algerian malacologist Paul Maurice Pallary (1869–1942), who collected holotype [extinct in the 1930s due to introduced Common Carp Cyprinus carpio]

Salmo pelagonicus Karaman 1938icus (L.), belonging to: Pelagonia, the geographical plain between Greece and Macedonia, where it occurs

Salmo pellegrini Werner 1931 in honor of French zoologist Jacques Pellegrin (1873–1944), who collected and studied the fishes of Morocco (where this trout occurs), and described S. pallaryi, also from Morocco, in 1924

Salmo peristericus Karaman 1938icus (L.), belonging to: Perister (or Baba) Mountain, Macedonia, which overlooks Prespa Lake and Brajcinska Reka creek, co-type localities

Salmo platycephalus Behnke 1968 flat-headed, from platýs (Gr. πλατύς), flat, and kephalḗ (Gr. κεφαλή), head, referring to its “broad, comparatively flat head”

Salmo rhodanensis Fowler 1974ensis, Latin suffix denoting place: Rhône River (Rhodanus in Greco-Roman geography), France, type locality [name coined by Roule (1923) but was not published in an available way]

Salmo rizeensis Turan, Kottelat & Engin 2010ensis, Latin suffix denoting place: Rize, Turkey (both city and province), where it is widely distributed

Salmo salar Linnaeus 1758 presumably from the Latin salire, to jump, referring to its ability to leap up waterfalls and other obstructions during its spawning run; another explanation is that it derives from sal, Latin for salt, referring to its marine life-history stage

Salmo schiefermuelleri Bloch 1784 in honor of Johann Ignaz Schiffermüller (1727–1806), Austrian theologian and naturalist, who provided holotype (note that Bloch misspelled the name)

Salmo stomachicus Günther 1866 stomachikós (Gr. στομαχικός), disordered in the stomach, or having a stomach disease, referring to its stomach membranes, which are “conspicuously thicker in this species than in its congeners”

Salmo taleri (Karaman 1933) in honor of Zdravko Taler (d. 1954), an amateur scientist who helped Karaman access remote sites to collect interesting trout populations in Montenegro, including holotype of this species

Salmo tigridis Turan, Kottelat & Bektaş 2011 genitive singular of Tigris, referring to Tigris River drainage, where type locality (Çatak Stream, Van Province, Turkey) is situated

Salmo trutta Linnaeus 1758 Medieval Latin for trout, its origins obscure, perhaps from tructa (L.), a kind of trout, in turn derived from tróktes (Gr. τρώκτης), gnawer or nibbler

Salmo trutta fario Linnaeus 1758 a teutonic word for “salmon-trout,” perhaps from the Old High German forhana, possibly a falsa lectio (false reading) of sario (typeset with a long s, which looks like an f); according to Ausonius (ca. 310–ca. 395), sario was a fish intermediate between salmo and salar

Salmo trutta lacustris Linnaeus 1758 Latin for of or belonging to a lake (lacustrine), referring to its occurrence in large cool lakes (compared with the anadromous nominate subspecies) [treated as a full species by some workers]

Salmo viridis Doadrio, Perea & Yahyaoui 2015 Latin for green, referring to its diagnostic greenish coloration on some parts of body and fins

Salmo visovacensis Taler 1950 ensis, Latin suffix denoting place: Visovac Lake, Croatia, type locality

Salmo zrmanjaensis Karaman 1938 ensis, Latin suffix denoting place: Zrmanja River near Obrovac and Zegar, Croatia, type locality

Salmo (subgenus Acantholingua) Hadzisce 1960 acanthus (L.), from ákantha (Gr. ἄκανθα), thorn; lingua (L.), tongue, referring to glossohyal teeth of S. ohridanus

Salmo ohridanus Steindachner 1892anus (L.), belonging to: Lake Ohrid, Albania and Macedonia, where it is endemic

Salmo (subgenus Salmothymus) Berg 1908 etymology not explained, probably a combination of Salmo and Thymallus, the latter genus at one time housing S. obtusirostris, and probably reflecting Berg’s belief that Salmothymus is a connecting link between the two genera

Salmo montenigrinus (Karaman 1933)inus (L.), belonging to: Montenegro, where type locality (Morača River at Titograd) is situated

Salmo obtusirostris (Heckel 1851) obtusus (L.), blunt; rostris, Neo-Latin scientific adjective of rostrum (L.), snout, referring to its “remarkably short, blunt and rounded” snout (translation, although snout is better described as elongate, not short)

Salmo obtusirostris krkensis (Karaman 1927)ensis, Latin suffix denoting place: Krka River at Knin, Croatia, where it is endemic

Salmo obtusirostris salonitana (Karaman 1927)ana (L.), belonging to: Solin (Salona in Latin), Dalmatia, Croatia, near where Jadro River (type locality) is situated

Salmo zetensis (Hadzisce 1960)ensis, Latin suffix denoting place: Zeta River, tributary to Lake Skadar, Montenegro, type locality

Salvelinus Richardson 1836 Latinization of säibling, an old German word for char (Richardson credited the name to Nilsson 1832, who used it for the group “Salvelini”)

Salvelinus albus Glubokovsky 1977 Latin for white, allusion not explained, presumably referring to white belly and/or pelvic and anal fins, whose first rays are white

Salvelinus alpinus (Linnaeus 1758) Latin for alpine or of the Alps, referring to its occurrence in alpine lakes

Salvelinus alpinus erythrinus (Georgi 1775)inus, Latin adjectival suffix: erythrós (Gr. ἐρυθρός), red, presumably referring to its brilliant red sides and ventral region

Salvelinus alpinus oquassa (Girard 1854) named for Lake Oquassa, one of the Rangeley Lakes of Maine, USA; according to local anglers, on or about 10 October this trout leaves the deep waters of Moosemegantic Lake to spawn in Kenebago, an inlet, and then ascends into Lake Oquassa, where it spends a month before returning to Moosemegantic until the next October

Salvelinus alpinus orientalis Kirillov 1972 Latin eastern, allusion not explained, perhaps referring to its distribution in eastern Yakutia (or Sakha Republic), Russia

Salvelinus andriashevi Berg 1948 in honor of Russian ichthyologist Anatoly Petrovich Andriashev (1910–2009), who collected holotype

Salvelinus boganidae Berg 1926 of Boganida Lake, Khatanga River basin, Sakha-Yakutia, Russia, type locality

Salvelinus confluentus (Suckley 1859) Latin for flowing together, referring to “confluent spots of dark brown, or black” on upper surface of body

Salvelinus curilus (Pallas 1814) Latinization of Kuriles, referring to its occurrence in rivers of the Kurile (or Kuril) Islands of the Russian Far East

Salvelinus czerskii Dryagin 1932 patronym not identified but probably in honor of Jan Czerski (1845–1892), Polish paleontologist, geologist, geographer and explorer of Siberia, where this species occurs

Salvelinus drjagini Logashev 1940 patronym not identified but clearly in honor of Petr Amphilokhovich Drjagin (1893–1977), ichthyologist, hydrobiologist, fisheries specialist, and a pioneer of fisheries research in Russia and management of inland waters

Salvelinus elgyticus Viktorovsky & Glubokovsky 1981icus (L.), belonging to: Elgygytkhyn Lake, Anadyr River basin, Chukotski Peninsula, Russia, type locality

Salvelinus evasus Freyhof & Kottelat 2005 past participle of the Latin verb evadere (escape, get away), referring to how, unlike other deepwater chars (e.g., S. neocomensis, S. profundus), this one has evaded extinction despite heavy pollution

Salvelinus faroensis Joensen & Tåning 1970ensis, Latin suffix denoting place: Faroe Islands, where it is endemic

Salvelinus gracillimus Regan 1909 superlative of gracilis (L.), slender, i.e., very slender; according to Regan (1911), its body is “more elongate than any other Char, the greatest depth being contained from five and a half to six and a half times in the length of the fish, measured to the base of the caudal fin”

Salvelinus gritzenkoi Vasil’eva & Stygar 2000 in honor of salmon biologist Oleg F. Gritsenko (also spelled Gritzenko), the “main organizer” of the 1999 expedition to the north Kuril Islands of Russia, when holotype was collected (and where this species is endemic)

Salvelinus inframundus Regan 1909 infra (L.), underneath or below; mundus (L.), world, i.e., underworld, referring to Hellyal Lake (now known as Heldale Water, Hoy Island, Scotland), type locality, derived from Hel (or Hella), the Norse goddess of the underworld

Salvelinus jacuticus Borisov 1935icus (L.), belonging to: Yakutia (Latinized a Jacutia), a large area in northern-central and eastern Siberia, where it occurs

Salvelinus killinensis (Günther 1866) ensis, Latin suffix denoting place: Loch Killin, Scotland, where it is endemic

Salvelinus krogiusae Glubokovsky & Chereshnev 2002 in honor of Russian ichthyologist Faina Vladimirovna Krogius (1902–1989), who worked for about 50 years on Dal’nee Lake, Kamchatka, on the biology of Oncorhynchus nerka and on lake ecology in general (also, she collected holotype in 1938)

Salvelinus kronocius Viktorovsky 1978icus (L.), belonging to: Lake Kronotskoye drainage, Kamchatka, Russia, where it is endemic

Salvelinus kuznetzovi Taranetz 1933 patronym not identified, probably in honor of Russian fisheries biologist Ivan Ivanovich Kuznetsov (1885–1962), who was working in Kamchatka (where this char occurs) at the time

Salvelinus lepechini (Gmelin 1789) in honor of Ivan Ivanovoich Lepechin (also spelled Lepyokhin, 1737–1802), Russian physician, naturalist and explorer, who first described this species, but apparently did not name it, in 1780

Salvelinus leucomaenis (Pallas 1814) artificially formed Latin adjective of leúkōma (Gr. λεύκωμα), a white spot, referring to white- or cream-colored spots on body

Salvelinus leucomaenis imbrius (Jordan & McGregor 1925) Latin for rainy, similar to S. l. pluvius, whose name means rainy, from a Japanese name that means “rain fish”

Salvelinus leucomaenis japonicus Oshima 1961icus (L.), belonging to: Japan, where it is endemic

Salvelinus leucomaenis pluvius (Hilgendorf 1876) Latin for rainy, named for Ameno-uwo, vernacular name for this fish in Japan, meaning “rain fish” or “rain charr,” referring to how they move from one shallow mountain stream to another during the rainy season

Salvelinus levanidovi Chereshnev, Skopets & Gudkov 1989 in honor of the late Russian ichthyologist and hydrobiologist Vladimir Yakovlevich Levanidov (1913–1981), “a leading expert on and student of the biology of salmonid ecosystems of the [Russian] Far East” (translation)

Salvelinus lonsdalii Regan 1909 in honor of Hugh Lowther, 5th Earl of Lonsdale (1857–1944), who “presented” the type specimens

Salvelinus mallochi Regan 1909 in honor of Peter Duncan Malloch (1853–1921), the “well-known naturalist of Perth” (Scotland), where this trout occurs (he was also a renowned angler and tackle maker)

Salvelinus malma (Walbaum 1792) vernacular name for this trout used in the Kamchatka Peninsula in the 16th century

Salvelinus malma anaktavukensis Morrow 1973ensis, Latin suffix denoting place: from the Eskimo name Anaktuvuk (meaning Pass), “the place of the caribou crossing,” referring to the approximate center of its east-west distribution and general area of its type locality in Alaska

Salvelinus malma lordii (Günther 1866) in honor of John Keast Lord (1819–1872), English veterinarian, journalist and naturalist to the British North American Boundary Commission, who presented holotype to the British Museum (Natural History)

Salvelinus malma miyabei Oshima 1938 patronym not identified, probably in honor of Japanese botanist-mycologist Kingo Miyabe (1876–1951)

Salvelinus maxillaris Regan 1909 Latin for of or belonging to the jaw, presumably referring to its maxillary extending to below posterior margin of eye in females and beyond in males

Salvelinus murta (Saemundsson 1908) local name for this charr along Lake Thingvalla, Iceland, where it is endemic

Salvelinus neiva Taranetz 1933 local name (possibly of Eskimo origin) for this char in the Russian Far East, where it occurs

Salvelinus neocomensis Freyhof & Kottelat 2005ensis, Latin suffix denoting place: Neocomus, Latin name of Neuchâtel, referring to Lake Neuchâtel, Switzerland, where this char was endemic [extinct due to eutrophication; last collected in 1904]

Salvelinus perisii (Günther 1865) of Llyn Peris (also known as Llanberris), a lake in Snowdonia, Wales, type locality [replacement name for Salmo cambricus Günther 1862, preoccupied by S. cambricus Donovan 1806, a synonym of S. trutta]

Salvelinus profundus (Schillinger 1901) Latin for deep, referring to its occurrence in deepwater habitat of Lake Constance in Germany, Switzerland and Austria [extinct since the late 1970s due to eutrophication]

Salvelinus salvelinoinsularis (Lönnberg 1900) Latin for of islands or insular, proposed as a subspecies of S. salvelinus (=alpinus) from Lake Ella, Bear Island, Norway

Salvelinus schmidti Viktorovsky 1978 patronym not identified, probably in honor of Russian ichthyologist Petr Yulievich Schmidt (1872–1949), a prominent researcher on fishes of the Russian Far East, where this species occurs

Salvelinus struanensis (Maitland 1881)ensis, Latin suffix denoting place, not explained in original description, but probably honors the chief of Clan Donnachaidh (traditionally known as Robertson of Struan, or Struan Robertson or simply Struan), in whose ancient territories Loch Rannoch, Pertshire, Scotland (type locality), lay

Salvelinus taimyricus Mikhin 1949icus (L.), belonging to: Lake Taimyr, Russia, type locality

Salvelinus taranetzi Kaganowsky 1955 in memory of Russian ichthyologist Anatoly Yakovlevich Taranetz (1910–1941), for significant contributions to the study of fishes from the Russian Far East and the genus Salvelinus

Salvelinus thingvallensis (Saemundsson 1908)ensis, Latin suffix denoting place: Lake Thingvalla, Iceland, where it is endemic

Salvelinus tolmachoffi Berg 1926 in honor of Innokenty Petrovich Tolmachov (1872–1950), geographer, geologist and paleontologist, who led an expedition to Khatanga, Russia, in 1905–06 and collected holotype

Salvelinus vasiljevae Safronov & Zvezdov 2005 in honor of ichthyologist Ekaterina Denisovna Vasil’yeva (b. 1952), for her significant contribution to the study of fishes from the Russian Far East, including Salvelinus

Salvelinus willoughbii (Günther 1862) in honor of Francis Willughby (sometimes spelled Willoughby, 1635–1672), British ichthyologist and ornithologist, “the first who with the practised eye of an ichthyologist examined the Charrs of England and Wales”

Salvelinus youngeri Friend 1956 in honor of Henry John Younger, of Younger’s Brewery (est. 1778). who gave permission for the “netting” in Loch Eck, Cowal, Argyllshire, Scotland, type locality

Salvelinus (subgenus Baione) DeKay 1842 baión (Gr. βαιόν), “a small fish alluded to by ancient writers,” referring to a spring population of dwarf or juvenile S. fontinalis that DeKay believed represented both a new genus and species (which he also named fontinalis, independent of S. fontinalis)

Salvelinus fontinalis (Mitchill 1814) Latin for of a spring or fountain; “He lives in running waters only,” Mitchill wrote, “and not in stagnant ponds; and, therefore, the lively streams, descending north and south from their sources on Long-Island [New York, USA], exactly suit the constitution of this fish”

Salvelinus fontinalis agassizii (Garman 1885) in honor of Swiss-born American zoologist-geologist Louis Agassiz (1807–1873), Harvard University, whose 1839 Histoire naturelle des poissons d’eau douce de l’Europe centrale is cited several times in Garman’s work, and who described S. namaycush siscowet in 1850 [extinct due to stocked gamefishes, last collected in 1930; treated as a full species by some workers]

Salvelinus fontinalis timagamiensis Henn & Rinkenbach 1925 ensis, Latin suffix denoting place: Timagami region of Ontario, Canada, type locality

Salvelinus (subgenus Cristivomer) Gill & Jordan 1878 crista (L.), crest; vomer (L.), plowshare, but in ichthyology the bone forming the front part of the roof of the mouth, referring to raised crest behind head of vomer, armed with teeth

Salvelinus namaycush (Walbaum 1792) Native American name for this species, probably the Cree namekush, meaning “dweller of the deep”

Salvelinus namaycush siscowet (Agassiz 1850) Native American name for this trout along Lake Superior in Ontario, Canada, derived from an Ojibwe word meaning “cooks itself” or “that which has oily flesh” (also apparently root word for “cisco,” vernacular name for Coregonus species in North America)

Salvethymus Chereshnev & Skopets 1990 combination of Salvelinus and Thymallus, reflecting how it possesses a number of morphological characteristics found in both genera

Salvethymus svetovidovi Chereshnev & Skopets 1990 in honor of Russian ichthyologist and Corresponding Member of the Academy of Sciences of the USSR, the late Anatoliy Nikolayevich Svetovidov (1903–1985), for his “considerable” (translation) contribution to the study of salmonids